Dryopteris

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Photo	Pteridium	Polystichum			(dryopteris)
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Dryopteris Adanson, 1763 Alvejuuret Wood Ferns Shield Ferns

Dryopteris, Den Virtuella Floran [DVF]

Dryopteris (Polypodiaceae); [TSF, 3]; [FPN, 50]

Foodplants for Blasticotoma filiceti filiceti, Petrophora chlorosata

Dryopteris filix-mas (L.) Schott, 1834 Kivikkoalvejuuri Male-fern Träjon Echter Wurmfarn Dryoptéris Fougère mâle
- Polypodium filix-mas L., 1753
- Aspidium filix-mas L. Swartz, 1801
- Dryopteris filix-mas; [TFG, 114]; [TSF, #11]; [FPN, 51]; [RK, 49]
783x1044 (~180Kb) Finland, Nokia, Utola, 20.6.2005, Photo © Harri Arkkio 902x640 (~118Kb) Finland, Nokia, Utola, 20.6.2005, Photo © Harri Arkkio
Dryopteris cristata (L.) Gray, 1848 Korpialvejuuri Gray Crested Shield Fern Narrow Swamp Fern Crested Wood Fern Granbräken Kammfarn Dryoptéris à crétes
- Polypodium cristatum L., 1753
- Aspidium cristatum L. Swartz, 1801
- Nephrodium cristatum L. Michaux, 1803
- Dryopteris cristata, Bilder ur Nordens Flora (in Swedish) [RUNEBERG]
- Dryopteris cristata; [TFG, 106]; [TSF, #12]; [FPN, 51]; [RK, 49]
Dryopteris clintoniana (Eaton) Dowell, 1906 Broad Swamp Fern Clinton's Wood Fern
- Aspidium cristatum v. clintonianum Eaton, 1867
- Dryopteris cristata v. clintoniana (Eaton) Underwood, 1893
- Dryopteris clintoniana; [TFG, 108]
Dryopteris celsa (Palmer) Small, 1958 Log Fern
- Dryopteris goldiana ssp. celsa Palmer, 1899
- Dryopteris celsa; [TFG, 110]
Dryopteris goldiana (Hooker) Gray, 1848 Giant Wood Fern Goldie's Wood Fern
- Aspidium goldianum Hooker, 1822
- Nephrodium goldianum Hooker Hooker & Greville, 1829
- Dryopteris goldiana; [TFG, 112]

Dryopteris carthusiana (Vill.) H.P. Fuchs Metsäalvejuuri Narrow Buckler-fern Spinulose Shield Fern Skogsbräken

Dryopteris spinulosa Roth Kunze
Dryopteris carthusiana [RK, 50]

700x571 (~104Kb) FINLAND: Ka: Virolahti, Virojoki 671:53, 19.9.1998, Photo © Markku Savela
Dryopteris carthusiana det. Tapio Lahtonen

900x601 (~79Kb) FINLAND: Ka: Virolahti, Virojoki 671:53, 19.9.1998, Photo © Markku Savela
Dryopteris carthusiana det. Tapio Lahtonen

700x583 (~38Kb) FINLAND: Ka: Virolahti, Virojoki 671:53, 19.9.1998, Photo © Markku Savela
Dryopteris carthusiana det. Tapio Lahtonen

Foodplant for Udea decrepitalis

? ssp. - Toothed Wood Fern Spinulose Wood Fern
- Dryopteris lanceolatocristata (Hoffmann) Alston, 1957
- Polypodium lanceolatocristata Hoffmann, 1790
- Polypodium spinulosum Mueller, 1767-77
- Aspidium spinulosum (Mueller) Swartz, 1801
- Dryopteris spinulosa (Mueller) Watt, 1867
- Dryopteris austriaca v. spinulosa (Mueller) Fiori, 1943
- Dryopteris spinulosa; Watt, 1867, [TFG, 120 (invalid?)]

Dryopteris carthusiana × cristata
- Dryopteris × uliginosa A. Braun ex Döll Druce
- Dryopteris carthusiana × cristata; [RK, 50]
Dryopteris expansa (C. Presl.) Fraser-Jenkins & Jermy Isoalvejuuri Nordbräken
- Dryopteris assimilis S. Walker
- Dryopteris austriaca auct.
- Dryopteris dilatata auct.
- Dryopteris expansa; [RK, 50]
Dryopteris dilatata (Hoffm.) A. Gray, 1848 Etelänalvejuuri Broad Buckler-fern Lundbräken
- Polypodium dilatatum Hoffmann, 1795
- Dryopteris austriaca auct.
- Dryopteris dilatata; [TFG, 118]; [RK, 50]
- v. (?) Mountain Wood Fern Spreading Wood Fern
  - Aspidium spinulosum v. anadenium Robinson, 1907
  - Dryopteris spinulosa v. americana (Fischer) Fernald, 1915
  - Aspidium spinulosa v. americanum Fischer, 1848
  - Dryopteris campyloptera (Kunze) Clarkson, 1930
  - Aspidium campylopterum Kunze, 1848
  - Dryopteris dilata; [TFG, 118]
Dryopteris austriaca (Jacq.) Woynar Stachliger Wurmfarn Dryoptéris d'Austriche Spreading Wood-fern
- Dryopteris spinulosum Sw.
- Dryopteris austriaca; [TSF, #14]
- Drypoteris austriaca; [FPN, 51]

Dryopteris fragrans (L.) Schott, 1834 Tuoksualvejuuri Fragrant Shield Fern Fragrant Wood Fern Doftbräken

Polypodium fragrans L., 1753

Dryopteris fragrans; [TFG, 104]; [RK, 51]

732x500 (~146Kb) Above the village Novyi Akatuy, the junction of the mountain ranges Uryumkanskii and Kukul'bei, Aleksandrovskozavodskoi District, E Chita Region, E Transbaikalia, Siberia, Russia. 23rd July 1997, Photo © Oleg Kosterin

v. remotiuscula (Komarov) Komarov, 1934
- Nephrodium fragrans v. remotiusculum Komarov, 1911
- Dryopteris fragrans v. remotiuscula; [TFG, 105 (note)]

Dryopteris villarsii (Bell.) Woynar Starrer Wurmfarn Dryoptéris de Villars Dryoptéris rigide
- Aspidium rigidum Sw.
- Dryopteris villarsii; [TSF, #13]
Dryopteris marginalis (L.) Gray, 1848 Marginal Wood Fern Evergreen Wood Fern Leather Wood Fern
- Polypodium marginalis L., 1753
- Aspidium marginale L. Swartz, 1801
- Nephrodium marginale L. Michaux, 1803
- Dryopteris marginalis; [TFG, 116]
Dryopteris intermedia (Willdenow) Gray, 1848 Fancy Fern Evergreen Wood Fern Intermediate Wood Fern
- Polypodium intermedium Muhlenberg, 1810
- Aspidium intermedium Willdenow, 1810
- Dryopteris spinulosa v. intermedia (Willdenow) Underwood, 1893
- Dryopteris spinulosa v. intermedia (Willdenow) Morton, 1950
- Dryopteris intermedia; [TFG, 122]
Dryopteris arguta (Kaulf.) Watt. Coastal Shield-fern
- Dryopteris arguta; [FPN, 51]

Other hybrids

Dryopteris celsa × cristata Wherry Dismal Swamp Fern
- Dryopteris × atroplaustris Small, 1938
- Dryopteris celsa × cristata; [TFG, 124]
Dryopteris celsa × intermedia Wherry Spaced Wood Fern
- Dryopteris × separabilis Small, 1938
- Dryopteris celsa × intermedia; [TFG, 124]
Dryopteris clintoniana × cristata Wherry, 1961 Deceptive Wood Fern
- Dryopteris clintoniana × cristata; [TFG, 126]
Dryopteris clintoniana × goldiana Dowell, 1908 Simulated Log Fern
- Dryopteris clintoniana × goldiana; [TFG, 126]
Dryopteris clintoniana × intermedia Dowell, 1908 Dowell's Wood Fern
- Filix dowellii Farwell, 1925
- Dryopteris × dowellii (Farwell) Wherry, 1961
- Dryopteris clintoniana × intermedia; [TFG, 126]
Dryopteris clintoniana × marginalis Slosson, 1910
- Dryopteris clintoniana × marginalis; [TFG, 126]
Dryopteris clintoniana × spinulosa Benedict, 1900 Benedict's Wood Fern
- Filix benedictii Farwell, 1923
- Dryopteris × benedictii (Farwell) Wherry, 1961
- Dryopteris clintoniana × spinulosa; [TFG, 126]
Dryopteris cristata × goldiana Benedict, 1909
- Dryopteris cristata × goldiana; [TFG, 128]
Dryopteris cristata × intermedia Dowell, 1908 Glandular Swamp Fern Boott's Wood Fern
- Aspidium boottii Tuckerman, 1843
- Dryopteris × boottii (Tuckerman) Underwood, 1893
- Dryopteris cristata × intermedia; [TFG, 128]
Dryopteris cristata × marginalis (Davenport, 1894) Miss Slosson's Wood Fern
- Aspidium cristatum × marginale Davenport, 1894
- Nephrodium slossonae Hahne, 1904
- Dryopteris × slossonae (Hahne) Wherry, 1942
- Dryopteris cristata × marginalis; [TFG, 128]
Dryopteris cristata × spinulosa Christensen, 1905 Braun's Wood Fern
- Aspidium spinulosum v. uliginosum Braun, 1843
- Dryopteris × uliginosa (Braun) Druce, 1909
- Dryopteris cristata × spinulosa; [TFG, 128]
Dryopteris filix-mas × goldiana Rugg, 1950
- Dryopteris "filix-mas" × goldiana; [TFG, 130]
Dryopteris filix-mas × marginalis Winslow, 1910 Vermont Wood Fern
- Dryopteris "filix-mas" × marginalis; [TFG, 130]
Dryopteris fragrans × intermedia
- Dryopteris fragranas v. remotiuscula × spinulosa v. intermedia Tryon, 1942
- Dryopteris fragrans × intermedia; [TFG, 130]
Dryopteris goldiana × intermedia Dowell, 1908
- Dryopteris goldiana × intermedia; [TFG, 130]
Dryopteris goldiana × marginalis Dowell, 1908 Susquehanna Wood Fern
- Dryopteris × leedsii Wherry, 1942
- Dryopteris goldiana × marginalis; [TFG, 130]
Dryopteris goliana × spinulosa Benedict, 1909 Poyser's Wood Fern
- Nephrodium cristatum ssp. clintonianum f. silvaticum Poyser, 1908
- Dryopteris × poyseri Wherry, 1960
- Dryopteris goliana × spinulosa; [TFG, 130]
Dryopteris intermedia × marginalis Benedict, 1909
- Dryopteris intermedia × marginalis; [TFG, 132]
Dryopteris intermedia × spinulosa Wherry, 1960 Triploid Wood Fern
- Dryopteris × triploidea Wherry, 1960
- Nephrodium spinulosum ssp. fructuosum Gilbert, 1901
- Dryopteris spinulosa v. fructuosa (Gilbert) Trudell, 1930
- Dryopteris intermedia f. fructuosa (Gilbert) Clute, 1938
- Dryopteris intermedia × spinulosa; [TFG, 132]
Dryopteris marginalis × spinulosa Slosson, 1904 Pittsford Wood Fern
- Dryopteris × pittsfordensis Slosson, 1904
- Dryopteris marginalis × spinulosa; [TFG, 132]

23.5.2018 (14)

References:

[DVF] Den Virtuella floran
http://linnaeus.nrm.se/flora/welcome.html; Ett projekt vid Naturhistoriska riksmuseet (Stockholm, Sweden).; Even for non-Swedish speaker, the distribution maps on species are nice (map buttons "Norden" and "Norra halvklotet")
[FPN] Hitchcock & Cronquist, 1998
Flora of the Pacific Northwest (11th ed.); ISBN 0-295-95273-3
[Harri Arkkio]
Finland; For the time being these pictures are only for this site. Other usage not allowed.
[Oleg Kosterin] Institue of Cytology & Genetics
Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[RK] Hämet-Ahti, L., Suominen, J., Ulvinen, T. & Uotila, P. (eds), 1998
Retkeilykasvio (Field Flora of Finland); Ed. 4. 656 pp.; Finnish Museum of Natural History, Botanical Museum. Helsinki
[RUNEBERG] Project Runeberg: Nordic literature online
Project Runeberg; Bilder ur Nordens Fauna (Carl Axel Magnus Lindman)
[TFG] Edgar T. Therry, ?1995
The Fern Guide: Northeasten and Midland United States and Adjacent Canada; [Original publication 1961]
[TSF] Eduard Thommen, 1951
Taschenatlas der Schweizer Flora

If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.