Ypthima

Ypthima Hübner, 1818 Rings Ringlets

Ypthima Hübner, 1818; Zuträge Samml. exot. Schmett. 1: 17 [missid. TS Papilio philomela Linnaeus]; TS: Ypthima huebneri Kirby
Xois Hewitson, 1865; Trans. ent. Soc. Lond. (3) 2 (4): 282; TS: Xois sesara Hewitson
Kolasa Moore, 1891; (nomen nudum)
Thymipa Moore, 1891; (nomen nudum)
Kolasa Moore, 1893; Lepidoptera Indica 2 (14): 57, (15):82; TS: Satyrus chenui[sic, recte chenu] Guérin-Méneville
Thymipa Moore, 1893; Lepidoptera Indica 2 (14): 57, 58; TS: Papilio baldus Fabricius
Nadiria Moore, 1893; Lepidoptera Indica 2 (14): 57, (15):85; TS: Ypthima bolanica Marshall
Pandima Moore, 1893; Lepidoptera Indica 2 (14): 58, (15):86; TS: Satyrus nareda Kollar
Lohana Moore, 1893; Lepidoptera Indica 2 (14): 58, (16):92; TS: Yphthima[sic] inica Hewitson
Dallacha Moore, 1893; Lepidoptera Indica 2 (14): 58, (16):94; TS: Yphthima[sic] hyagriva Moore
Shania Evans, 1912; J. Bombay nat. Hist. Soc. 21 (2): 564; TS: Ypthima megalia de Nicéville
Sundaypthima Uemura, 1996

Iphthima; Westwood, [1851], Gen. diurn. Lep. (2): pl. 67 (missp.)
Yphthima; Scudder, 1875, Proc. Amer. Acad. Arts Sci. 10 (2): 289 (missp.)
Thympia; Hemming, 1967, Bull. Br. Mus. (Ent.) Suppl. 9: 442 (missp.)
Ypthima (Satyrinae); Moore, [1880], Lepid. Ceylon 1 (1): 24
Ypthima (Satyrina); [Wahlberg]
Ypthima (Ypthimina); Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 183; [AfrL]
Ypthima (Ypthimini); [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 43

816x612 (~79Kb) upperside Thailand, Chantaburi Province, Khao-Khitchakut National Park, a floating bog on a bank of the pond at headquarters. 30th January 2005, Photo © Oleg Kosterin	743x786 (~67Kb) underside Thailand, Chantaburi Province, Khao-Khitchakut National Park, a floating bog on a bank of the pond at headquarters. 30th January 2005, Photo © Oleg Kosterin	600x400 (~50Kb) underside India: Melghat, 9.10.2008, Photo © Kedar Tokekar
928x820 (~101Kb) underside Thailand, Chantaburi Province, Khao-Khitchakut National Park, a bank of the pond at headquarters. 7th January 2006, Photo © Oleg Kosterin	512x341 (~35Kb) upperside India: Guwahati, 17.11.2005, Photo © Kedar Tokekar	512x341 (~36Kb) India: Ziro, 9.11.2005, Photo © Kedar Tokekar
1137x989 (~250Kb) underside male female Thailand, Phuket, a garden in the Nai Yang beach area. 16th February 2009, Photo © Oleg Kosterin	810x827 (~203Kb) underside Thailand, Phuket, a bank of a pond at a road 1 km NE of Khao Sai Tan Kliang Mt., 18th February 2009, Photo © Oleg Kosterin	1066x955 (~257Kb) underside Cambodia, Kep Province, Kep. 22th April 2010, Photo © Oleg Kosterin
846x819 (~194Kb) underside Cambodia, Kep Province, Kep, Treetop Buingalow. 6th December 2010, Photo © Oleg Kosterin

The exact identification of these species is still unknown, but tentatively assumed to belong into this group.

Subgenus Thymipa Moore, 1893

Thymipa Moore, 1893; Lepidoptera Indica 2 (14): 57, 58; TS: Papilio baldus Fabricius

Thymipa; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

The sakra species-group

Sakra; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima similis Elwes & Edwards, 1893

Assam, Manipur, Burma, Thailand, Laos. See [maps] Eastern Five-Ring

Ypthima similis Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1): 29, pl. 1, f. 19; TL: Karen Hills
yoma Evans, 1924²; J. Bombay nat. Hist. Soc. 29 (3): 787; TL: Pegu
- = ; [NHM card]

Ypthima similis, Butterflies in Indo-China [Yutaka Inayoshi]

Ypthima similis; [BOR], 465
Ypthima (Thymipa) similis; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

861x909 (~89Kb) underside Thailand, Chon Buri Province, Pattaya, a vasting land with open tree stand behind the Jomtien beach, 1st February, 2005, Photo © Oleg Kosterin

684x754 (~82Kb) upperside Thailand, Chon Buri Province, Pattaya, Ko Khram Island, Saikaew beach, a bank of a pond in a wood near the coast, 3rd February, 2006, Photo © Oleg Kosterin

Y. s. similis S.Yunnan, ...
- Ypthima similis similis; Huang & Xue, 2004, Neue Ent. Nachr. 57: 141 (name), pl. 11, f. 7

Ypthima iarba de Nicéville, 1895 -. See [maps]
- Ypthima iarba de Nicéville, 1895; J. Bombay nat. Hist. Soc. 10 (1): 18, pl. R, f. 7-8; TL: Sumatra
- Ypthima iarba, Butterflies in Indo-China [Yutaka Inayoshi]
- Ypthima jarbas[sic]; [NHM card]
- Ypthima iarba; [BOR], 472
- Ypthima (Thymipa) iarba; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Y. i. iarba Burma, Sumatra, ?Peninsular Malaya
  - Ypthima iarba iarba; [BOR], 472
- Y. i. gaugamela Fruhstorfer, 1911² E.Java
  - iarba gaugamela Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: pl. 99 d
  - Ypthima iarba gaugamela; [BOR], 472
  - Ypthima (Thymipa) gaugamela; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Y. i. eupeithes Fruhstorfer, 1911² W.Java
  - iarba eupeithes Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 287, pl. 99 f
  - Ypthima iarba eupeithes; [BOR], 472
  - Ypthima (Thymipa) eupeithes; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima affectata Elwes & Edwards, 1893 Assam, Manipur. See [maps]
- Ypthima affectata Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1): 30, pl. 1, f. 30; TL: Terria Ghat; Mamloo, Khasias
- Ypthima affectata, Butterflies in Indo-China [Yutaka Inayoshi]
- Ypthima affectata; [BOR], 465
- Ypthima (Thymipa) affectata; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima sobrina Elwes & Edwards, 1893 Assam, Burma, Thailand, Laos, S.Yunnan. See [maps]
- Ypthima sobrina Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1): 29, pl. 1, f. 17-18; TL: Karen Hills
- Ypthima sobrina, Butterflies in Indo-China [Yutaka Inayoshi]
- Ypthima sobrina; [BOR], 465
- Ypthima (Thymipa) sobrina; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Ypthima sobrina; Huang & Xue, 2004, Neue Ent. Nachr. 57: 141 (name)
Ypthima savara Grose-Smith, 1887 Manipur - Burma - Indo China. See [maps] Pallid Five-Ring
- Yphthima[sic] savara Grose-Smith, 1887; Ann. Mag. nat. Hist. (5) 20: 267; TL: Burma-Siam frontier
- Ypthima savara savara, Butterflies in Indo-China [Yutaka Inayoshi]
- Ypthima savara; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 37, pl. 2, f. 30; [BOW]: pl. 173, f. 8; [BOR], 471
- Ypthima (Thymipa) savara; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Y. s. savara Burma, Thailand, Laos, S.Yunnan
  - Ypthima savara savara; Huang & Xue, 2004, Neue Ent. Nachr. 57: 141 (name), pl. 11, f. 6
- Y. s. tonkiniana Fruhstorfer, 1908 N.Vietnam
  - Ypthima ? tonkiniana Fruhstorfer, 1908; Ent. Zs. 22 (31): 127; TL: Tonkin, Chiem-Hoa
  - Ypthima savara tonkiniana Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9: 292, pl. 99, f. 2; TL: Chiem Hoa, N.Vietnam
  - Ypthima savara tonkiniana, Butterflies in Indo-China [Yutaka Inayoshi]
  - Ypthima savara tonkiniana; [BMP]: 136, pl. 17, f. 20-21
Ypthima sakra Moore, 1857 Murree - Assam, Karen Hills. See [maps] Himalayan Five-Ring
- Yphthima[sic] sakra Moore, 1857; in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1: 236
- Yphthima[sic] sakra; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4): 290, pl. 18, f. 18
- Ypthima sakra; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 40; [BOW]: pl. 173, f. 9; [BOR], 472; [MRS], 392
- Ypthima (Thymipa) sakra; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
512x341 (~37Kb) upperside India: Guwahati, 17.11.2005, Photo © Kedar Tokekar

Larva on Gramineae [MRS]
- Y. s. sakra Sikkim, Bhutan
  - Ypthima sakra sakra; [BOR], 472
- Y. s. austeni (Moore, 1893) Assam, N.Burma, SE.Tibet (Yigong), Gooligonshan Mts.
  - Thymipa austeni Moore, 1893; Lepidoptera Indica 2 (15): 69, pl. 109, f. 3, 3a
  - Ypthima sakra austeni, Butterflies in Indo-China [Yutaka Inayoshi]
  - Ypthima austeni; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 51
  - Ypthima sakra austeni; [BOR], 472; Huang, 2001; Huang, 2003, Neue Ent. Nachr. 55: 98 (note)
- Y. s. nujiangensis Huang, 2001 SE.Tibet
  - Ypthima sakra nujiangensis Huang, 2001; Neue Ent. Nachr. 51: 94, pl. 7, f. 51; TL: Longpo to Nidadan
  - Ypthima sakra nujiangensis; Huang, 2003, Neue Ent. Nachr. 55: 98 (note)
- Y. s. leechi Forster, [1948] Sichuan
  - Ypthima sakra leechi Forster, [1948]; Mitt. Münch. Ent. Ges. 34 (2): 483, pl. 30-31, f. 16
  - Ypthima sakra leechi, Butterflies in Indo-China [Yutaka Inayoshi]
  - Ypthima sakra leechi; Huang, 2001, Neue Ent. Nachr. 51: 94 (note)
- Y. s. matinia Fruhstorfer, 1911² NW.India
  - sakra matinia Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 292
  - Ypthima sakra matinia; [BOR], 472
Ypthima pseudosavara Uémura & Monastyrskii, 2000 Vietnam. See [maps]
- Ypthima pseudosavara Uémura & Monastyrskii, 2000; Trans. Lepid. Soc. Japan 51 (2): 153, f. 1d-e, 3, 10-13; TL: Vietnam, Thanh Hoa Prov., Nhu Xuan, Ben En Nat. Park (105°21'-105°35'E, 19°30'-19°40'N)
- Ypthima pseudosavara, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima nikaea Moore, [1875] W.Himalayas, Nepal. See [maps]
- Yphthima[sic] nikaea Moore, [1875]; Proc. zool. Soc. Lond. 1874 (4): 567; TL: NW.Himalayas
  - = Ypthima sakra ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 40
  - = ?; Huang, 2001, Neue Ent. Nachr. 51: 94
- Ypthima nikaea; [BOR], 471; [MRS], 392
- Ypthima (Thymipa) nikaea; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima hannyngtoni Eliot, 1967 NW.India. See [maps]
- Ypthima hannyngtoni Eliot, 1967; Entomologist 100 (1246): 54; TL: Masuri
- Ypthima (Thymipa) hannyngtoni; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima methorina Oberthür, 1891 China (Kweichow, Ta-tsien-lo). See [maps]
- Ypthima methorina Oberthür, 1891; Étud. d'Ent. 15: 15, pl. 2, f. 15
- Ypthima methorina completa Oberthür, 1911; Étud. Lépid. Comp. 5 (1): 317, pl. 60, f. 551; TL: Thibet
  - = ; [NHM card]
- Ypthima methorina; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 484; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 39, pl. 2, f. 33; [BOW]: pl. 173, f. 9 (text only)
- Ypthima (Thymipa) methorina; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima medusa Leech, [1892] Szechwan. See [maps]
- Ypthima medusa Leech, [1892]; Butts China Japan Corea (1): 84, pl. 10, f. 6; TL: Szechwan
- Ypthima medusa; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 485; [MRS], 392
- Ypthima (Thymipa) medusa; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Larva on Gramineae [MRS]
Ypthima davidsoni Eliot, 1967 Assam. See [maps]
- Ypthima davidsoni Eliot, 1967; Entomologist 100 (1246): 54; TL: Assam, Khasias
- Ypthima (Thymipa) davidsoni; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima formosana Fruhstorfer, 1908 Taiwan
- Ypthima methora formosana Fruhstorfer, 1908; Ent. Zs. 22 (29): 119
- Ypthima conjuncta formosana; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 481; [BOR], 471
- Ypthima formosana; [MRS], 391
- Ypthima (Thymipa) formosana; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Ypthima formosana; Takahashi, 2000, Trans. lepid. Soc. Japan 51 (1): 2
Larva on Brachiaria mutica, Imperata cylindrica, Cynodon dactylon, Miscanthus sinensis, M. floridulus, Setaria palmifolia, Isachne globosa [MRS]
Ypthima atra Cantlie & Norman, 1959
- atra Cantlie & Norman, 1959; J. Bombay nat. Hist. Soc. 56 (1): 69
- Ypthima atra, Butterflies in Indo-China [Yutaka Inayoshi]
- Ypthima atra; [NHM card]
- Ypthima (Thymipa) atra; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima persimilis Elwes & Edwards, 1893 Bhutan, Assam, Manipur. See [maps]
- Ypthima persimilis Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1): 39
- Ypthima persimilis, Butterflies in Indo-China [Yutaka Inayoshi]
- Ypthima persimilis; [BOR], 470
- Ypthima (Thymipa) persimilis; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima evansi Eliot, 1967 Thailand, Burma. See [maps]
- Ypthima evansi Eliot, 1967; Entomologist 100 (1246): 56; TL: North Shan States
- Ypthima evansi evansi, Butterflies in Indo-China [Yutaka Inayoshi]
- Ypthima (Thymipa) evansi; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Y. e. haka Eliot, 1967 Burma
  - Ypthima evansi haka Eliot, 1967; Entomologist 100 (1246): 58; TL: Burma North Chin Hills
Ypthima dohertyi (Moore, [1893]) Manipur - S.Burma. See [maps] Great Five-Ring
- Thymipa dohertyi Moore, [1893]; Lepidoptera Indica 2 (15): 65, pl. 108, f. 1, 1a; TL: Upper Tenasserim
- Ypthima dohertyi dohertyi, Butterflies in Indo-China [Yutaka Inayoshi]
- Ypthima dohertyi; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 38, pl. 2, f. 31; [BOR], 470
- Ypthima (Thymipa) dohertyi; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Y. d. dohertyi Burma, Thailan, Laos
- Y. d. khasia Eliot, 1967 Assam
  - Ypthima dohertyi khasia Eliot, 1967; Entomologist 100 (1246): 58; TL: Assam, Khasia
- Y. d. mossmani Eliot, 1967
  - Ypthima dohertyi mossmani Eliot, 1967; Entomologist 100 (1246): 59, pl. 2
  - Ypthima dohertyi mossmani, Butterflies in Indo-China [Yutaka Inayoshi]
  - Ypthima dohertyi mossmani; [BMP]: 136, pl. 17, f. 22
Ypthima methora Hewitson, 1865 Sikkim - Assam, N.Burma, Bhutan, Nepal. See [maps] Variegated Five-Ring
- Yphthima[sic] methora Hewitson, 1865; Trans. ent. Soc. Lond. (3) 2 (4): 291, pl. 18, f. 20-21; TL: N.India
- gela Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 292
  - = ; [NHM card]
- Ypthima methora; Moore, 1878, Proc. zool. Soc. Lond. 1878 (4): 825; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 37, pl. 2, f. 29; [BOW]: pl. 173, f. 3; [BOR], 470; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Y. m. methora Nepal, ...
  - Ypthima methora methora; Morishita & Inomata, 1998, Trans. lepid. Soc. Japan 49 (1): 3
Ypthima conjuncta Leech, 1891 W.China, C.China, Taiwan. See [maps]
- Ypthima conjuncta Leech, 1891; Entomologist 24 (Suppl.): 66; TL: Ta-chien-lu, Moupin; Chia-Kow-Ho, Omei-Shan; Wa-shan; huang-mu-chang; Chang-Yang; Ichang
- Ypthima conjuncta luxurians Forster, [1948]; Mitt. Münch. Ent. Ges. 34 (2): 482, pl. 32-33, f. 1-2; TL: Fukien, Kuatun, 2300m
  - = ; [NHM card]
- Ypthima conjuncta; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 39, pl. 2, f. 32; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 481; [BOW]: pl. 205, f. 1; [BOR], 471; [MRS], 391
- Ypthima (Thymipa) conjuncta; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Y. c. conjuncta C.China, W.China
  - Ypthima conjuncta conjuncta; [BOR], 471
- Y. c. monticola Uemura & Koiwaya, 2000 Yunnan
  - Ypthima conjuncta monticola, Butterflies in Indo-China [Yutaka Inayoshi]
  - Ypthima conjuncta monticola; Huang, 2003, Neue Ent. Nachr. 55: 100 (note)
Ypthima microphthalma Forster, [1948]
- Ypthima methora microphthalma Forster, [1948]; Mitt. Münch. Ent. Ges. 34 (2): 484, pl. 32-33, f. 3; TL: N.Yunnan, Likiang, Yülingshan, ca. 3000m
- Ypthima microphthalma; Huang & Wu, 2003, Neue Ent. Nachr. 55: 122 (note)
- Ypthima (Thymipa) microphthalma; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

The tappana species-group

Tappana; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima yayeyamana Nire, 1920 Japan (Iriomotejima, Ishigakijima). See [maps]
- Ypthima (Thymipa) yayeyamana; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima tappana Matsumura, 1909 Taiwan. See [maps]
- Ypthima tappana Matsumura, 1909; Ent. Zs. 23 (19): 92; TL: Formosa, Tappan, 4000ft
- Ypthima tappana; [BOR], 472 (note); [MRS], 392; Uemura, 1984, Tyô to Ga 35 (3): 137
- Ypthima (Thymipa) tappana; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Y. t. continentalis Murayama, 1981 Yunnan, Sichuan
  - Ypthima tappana continentalis Murayama, 1981; ; TL: Sichuan, Qingchenshan
  - Ypthima tappana continentalis; Huang, 2003, Neue Ent. Nachr. 55: 100 (note)
Ypthima cantlei Norman, 1958 Assam (Kangpokpi). See [maps]
- cantlei Norman, 1958; J. Bombay nat. Hist. Soc. 55 (1): 180; TL: Assam
- Ypthima cantliei[sic]; [NHM card]
- Ypthima cantlei; [BOR], 465
- Ypthima (Thymipa) cantlei; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

The philomela species-group

Philomela; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima philomela (Linnaeus, 1763) India (Nilgiri-Wynaad), N.Burma, ?Peninsular Malaya, Java, Sumatra. See [maps] Baby Fourring
- Papilio philomela Linnaeus, 1763; Amoenitates Acad. 6: 404; TL: Java
  - = ?Ypthima baldus ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 14
- Ypthima laroides Westwood, [1851]; Gen. diurn. Lep. (2): 395
  - = ; [NHM card]
- Ypthima tabella Marshall & de Nicéville, 1883; Butts India Burmah Ceylon 1 (2): 234
  - = Ypthima baldus ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 14
  - = ; [NHM card]
- peguana Evans, 1924²; J. Bombay nat. Hist. Soc. 29 (3): 786; TL: N.Burma
  - = ; [NHM card]
- Ypthima philomela; [BOW]: pl. 173, f. 6; [BOR], 465
- Ypthima (Thymipa) philomela; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima baldus (Fabricius, 1775)

Himalayas (Chamba - Assam), Burma, India, Thailand, China, Korea, Japan, Ussuri, Kuriles. See [maps] Common Five-Ring

Papilio baldus Fabricius, 1775; Syst. Ent.: 829, no. 202-3; TL: "India"
Ypthima argus Butler, 1866; J. Linn. Soc. Zool. Lond. 9 (34): 56; TL: Japan, Hakodate
Ypthima prattii Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1): 35, pl. 3, f. 55; TL: Ichang
- = Yptihma argus ; [NHM card]
Ypthima baldus ? ishigakina Fruhstorfer, 1908; Ent. Zs. 22 (31): 127; TL: Ishigaki
- = ; [NHM card]
gallienus Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 288; TL: Hainan
- = ; [NHM card]
scota Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 289; TL: Hongkong
- = ; [NHM card]
morus Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 289; TL: Sumatra
- = ; [NHM card]
satpura Evans, 1924²; J. Bombay nat. Hist. Soc. 29 (3): 787
- = ; [NHM card]
madrasa Evans, 1924²; J. Bombay nat. Hist. Soc. 29 (3): 787; TL: S.India
- = ; [NHM card]
Ypthima argus ab. narensis Sugitani, 1932; Zephyrus 4 (?): 90; TL: Japan
- = Yptihma argus ; [NHM card]
Ypthima baldus kangeana Kalis, 1933; Tijdschr. Ent. 76 (1-2): 55; TL: Kangean
- = ; [NHM card]

Ypthima baldus, Lepiforum [LEPIFORUM]
Ypthima baldus baldus, Butterflies in Indo-China [Yutaka Inayoshi]

Yphthima[sic] baldus; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4): 286
Ypthima argus; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 35, pl. 2, f. 28; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 476; Kudrna, 1974, Atalanta 5: 99; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 43
Ypthima (Thymipa) argus; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima baldus; Moore, 1878, Proc. zool. Soc. Lond. 1878 (4): 825; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 14, pl. 1, f. 15-16; [BRU], 192; [BOW]: pl. 172, f. 15; [BOR], 466
Ypthima balda; [MRS], 390
Ypthima (Thymipa) baldus; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

664x564 (~92Kb) Korea, 1989, S.W. Cheong col. & det., Photo © Enrique Garcia-Barros	1000x699 (~120Kb) underside An edge of a shady bamboo forest at Dragonfly Biotope Ponds, Hiraoka, Ikazaki-cho, Ehime-ken, Shikoku-Chiho, Japan. 7th August 2002 Photo © Oleg Kosterin	740x572 (~69Kb) upperside Japan: Tokyo, 1.9.2007, Photo © Peter Payzant
760x893 (~82Kb) underside Japan: Tokyo, 1.9.2007, Photo © Peter Payzant	512x341 (~33Kb) upperside India: Guwahati, 17.11.2005, Photo © Kedar Tokekar	512x341 (~41Kb) underside India: Guwahati, 17.11.2005, Photo © Kedar Tokekar

Larva on Poa annua, Echinochloa crusgalli, Digitaria , Elytrigia , (in Japan) [BRU], Fukuda et al., 1983 Microstegium ciliatum, Pogonatherum crinitinum [MRS]

Y. b. baldus India - Indo China, Burma (Tenasserim), Thailand, S.Yunnan
- Ypthima baldus baldus; [BOR], 466
Y. b. newboldi Distant, 1882 Peninsular Malaya, Langkawi Is., Singapore
- Ypthima newboldi Distant, 1882; Ann. Mag. nat. Hist. (5) 9 (53): 396; TL: Prov. Wellesley, Perak, Malaysia
  - = Ypthima marshalli ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 26
- Ypthima baldus newboldi, Butterflies in Indo-China [Yutaka Inayoshi]
- Ypthima baldus newboldi; [BMP]: 135, pl. 17, f. 25-27; [BOR], 466
Y. b. luoi Huang, 1999 Yunnan
- Ypthima baldus luoi; Huang, 2003, Neue Ent. Nachr. 55: 98 (note)
Y. b. moerus Fruhstorfer² Sumatra
- Ypthima baldus moerus; [BOR], 466
Y. b. selinuntius Fruhstorfer, 1911² Borneo, Natuna Is.
- baldus selinuntius Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 289, pl. 99 d; TL: Natuna Is.
- Ypthima baldus selinuntius; [BOR], 466
Y. b. pasitelides Fruhstorfer, 1911² Bawean
- baldus pasitelides Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 289; TL: Bawean
- Ypthima baldus pasitelides; [BOR], 466
Y. b. zodina Fruhstorfer, 1911 Taiwan
- baldus zodina (Fruhstorfer, 1911); in Seitz, Gross-Schmett. Erde 9: 289
- Ypthima baldus zodina; [BOR], 466
- Ypthima balda zodina; [MRS], 390
Larva on Miscanthus sinensis, M. floridulum, Isachne globosa, Brachiaria mutica, Oplismenus undulatifolius, Microstegium ciliatum, Pogonatherum crinitum [MRS]
Y. b. marshalli Butler, 1882
- Ypthima marshalli Butler, 1882; Ann. Mag. Nat. Hist. (5) 10: 373; TL: Thaungyin Valley, Meplay
- ?Ypthima evanescens Butler, 1881; Ann. Mag. Nat. Hist. (5) 7 (38): 134
  - = Ypthima marshalli ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 26
  - = Ypthima argus ; [NHM card]
- Ypthima marshalli; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 25, pl. 1, f. 1, 1a, 11
- Ypthima baldus marshalli; [MRS], 390
Y. b. hyampeia Fruhstorfer, 1911² S.Ussuri, Korea
- baldus hyampeia Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 290, pl. 99 e; TL: Amur
  - = Ypthima argus ; [NHM card]
- ?hampeia (Seitz, 1930); Gross-Schmett. Erde 1 (Suppl.): 131
  - = Ypthima argus ; [NHM card]
- Ypthima argus hyampeia; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 476, pl. 30-31, f. 7-8; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 43
Y. b. jezoensis Matsumura, 1919 Kuriles
- Ypthima argus var. jezoensis Matsumura, 1919; Thous. Ins. Japan. Addit. 3: 728; TL: Hokkaido
  - = Ypthima argus argus ; Uemura, 1984, Tyô to Ga 35 (3): 136
  - = Ypthima argus ; [NHM card]
- ?Ypthima argus jezoensis; Matsumura, 1928, Insecta Matsumurana 2 (4): 191
- Ypthima argus jesoensis[sic, recte jezoensis]; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 43
Y. b. okurai Okano, 1962 Formosa
- Ypthima argus okurai Okano, 1962; Tyô to Ga 12 (3): 52; TL: Sankakuhò (ca. 2370m), Central Formosa

Ypthima zodia Butler, 1871 Hainan. See [maps]
- Yphthima zodia Butler, 1871; Trans. ent. Soc. Lond. 1871 (3): 402
- zodiaca Moore, 1878²; Proc. zool. Soc. Lond. 1878 (3): 696
  - = Ypthima zodiaca ; [NHM card]
- Ypthima albescens Poujade, 1885; Bull. Soc. Ent. Fr. (6) 5: xli
  - = ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 28
- Ypthima zodia tapaishani Forster, [1948]; Mitt. Münch. Ent. Ges. 34 (2): 474, pl. 30-31, f. 2-3; TL: Tapaishan
  - = Ypthima zodiaca ; [NHM card]
- Ypthima zodia septentrionalis Forster, [1948]; Mitt. Münch. Ent. Ges. 34 (2): 474, pl. 30-31, f. 4; TL: Mienshan
  - = Ypthima zodiaca ; [NHM card]
- Ypthima melli Forster, [1948]; Mitt. Münch. Ent. Ges. 34 (2): 475, pl. 30-31, f. 10-11; TL: N.Yunna, Likiang
  - = Ypthima zodiaca ; [NHM card]
- pseudodromon Forster, [1948]; Mitt. Münch. Ent. Ges. 34 (2): 488
  - = Ypthima zodiaca ; [NHM card]
- Ypthima zodia albescens; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 473
- ?Ypthima (Thymipa) melli; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Ypthima zodia; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 27, pl. 1, f. 12-13; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 473, pl. 30, 31, f. 1; [MRS], 391
- Ypthima (Thymipa) zodia; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima horsfieldii Moore, 1884 -. See [maps]
- Ypthima horsfieldii Moore, 1884; J. asiat. Soc. Bengal 53 Pt.II (1): 18; TL: Java
- Ypthima horsfieldii; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 28
- Ypthima (Thymipa) horsfieldi; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Y. h. horsfieldii Java
  - ?Ypthima baldus horsfieldi; [BOR], 466
- Y. h. humei Elwes & Edwards, 1893
  - Ypthima humei Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1): 32, pl. 3, f. 46
  - Ypthima horsfieldii humei; [BMP]: 136, pl. 17, f. 28-29
Ypthima cerealis Watson, 1897 -. See [maps]
- Ypthima cerealis Watson, 1897; J. Bombay nat. Hist. Soc. 10 (4): 646, pl. A, f. 3-4; TL: Myingyan, Upper Burma
- Ypthima (Thymipa) cerealis; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Y. c. cerealis Burma, Thailand, Laos
- Y. c. singorensis Aoki & Uémura, 1984 Thailand
  - Ypthima cerealis singorensis Aoki & Uémura, 1984; Tokurana 6/7: 79, f. 2j, 6, 8B, 26, pl. 5, f. 3, 7; TL: Thailand
Ypthima nebulosa Aoki & Uémura, 1982 Sumatra, Thailand, Laos. See [maps]
- Ypthima nebulosa Aoki & Uémura, 1982; Memoirs of the Tsukada Coll. No. 4: 13, pl. 5, f. 22-23, pl. 6, f. 22-23; TL: Sumatra
- Ypthima nebulosa, Butterflies in Indo-China [Yutaka Inayoshi]
- Ypthima (Thymipa) nebulosa; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima yunosukei Aoki & Uémura, 1984 Thailand. See [maps]
- Ypthima yunosukei Aoki & Uémura, 1984; Tokurana (6/7): 81-82, pl. 5, f. 1-2, 5-6; TL: Phukadung, Thailand
- Ypthima yunosukei, Butterflies in Indo-China [Yutaka Inayoshi]
- Ypthima (Thymipa) yunosukei; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima fasciata Hewitson, 1865 Malaysia, Sumatra. See [maps]
- Yphthima[sic] fasciata Hewitson, 1865; Trans. ent. Soc. Lond. (3) 2 (4): 287; TL: Sarawak, Sumatra
- Ypthima fasciata; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 44; [BOW]: pl. 173, f. 1; [BOR], 468
- Ypthima (Thymipa) fasciata; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Y. f. fasciata Borneo, Natuna Is.
  - Ypthima fasciata fasciata; [BOR], 468
- Y. f. torone Fruhstorfer, 1911 Peninsular Malaya, Sumatra
  - Ypthima fasciata torone Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9: 288, (75) ; TL: Padang, Sumatra
  - Ypthima fasciata torone, Butterflies in Indo-China [Yutaka Inayoshi]
  - Ypthima fasciata torone; [BOR], 468; [BMP]: 136, pl. 17, f. 23-24
Ypthima indecora Moore, 1882 Kashmir - Kumaon. See [maps] Western Five-Ring
- Ypthima indecora Moore, 1882; Proc. zool. Soc. Lond. 1882 (1): 238, pl. 12, f. 7
- Ypthima indecora; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 13, pl. 1, f. 10, 14; [BOR], 465
- Ypthima (Thymipa) indecora; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima nynias Fruhstorfer, 1911² Sulawesi. See [maps]
- nigricans nynias Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 290
- Ypthima baldus nynias; [BOR], 466
- Ypthima (Thymipa) nynias; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 183
- Y. n. nynias Sulawesi, Muna, Buton
  - Ypthima (Thymipa) nynias nynias; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 183
- Y. n. aretas Fruhstorfer, 1911² Salayar
  - nynias aretas Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 290; TL: Saleyer
  - Ypthima baldus aretas; [BOR], 466
  - Ypthima (Thymipa) nynias aretas; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 183
- Y. n. gadames Fruhstorfer, 1911² Banggai
  - nynias gadames Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 290; TL: Bangkai
  - Ypthima gadames; [BOR], 472 (note)
  - Ypthima (Thymipa) nynias gadames; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 183
Ypthima stellera (Eschscholtz, 1821) Philippines. See [maps]
- Hipparchia stellera Eschscholtz, 1821; in Kotzebue, Entdeck. Reise Süd-See 3 (App. 5): 216, pl. 10, f. 24
- galeria Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 290; TL: Palawan
  - = ; [NHM card]
- Yphthima[sic] stellera; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4): 285
- Ypthima stellera; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 26, pl. 3, f. 45; [BOR], 472 (note)
- ?Ypthima (Thymipa) sterella[sic]; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Y. s. sepyra Hewitson, 1865 Bachan, Ternate, Halmahera
  - Yphthima[sic] sepyra Hewitson, 1865; Trans. ent. Soc. Lond. (3) 2 (4): 292, pl. 17, f. 10; TL: Gilolo, Batchian
    
    = Ypthima sepyra ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 27
  - Ypthima stellera sepyra; [BAUR], 265 (text)
Ypthima aphnius (Godart, [1824]) Celebes, Timor. See [maps]
- Satyrus aphnius Godart, [1824]; Encyclopédie Méthodique 9 (2): 551; TL: Timor
- Yphthima leuce Doherty, 1891; J. Asiat. Soc. Bengal 60 Pt.II (2): 169; TL: Sumba; Sumbawa
  - = ; [NHM card]
- caratonus Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 289; TL: Bali
  - = ; [NHM card]
- saravus Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 290; TL: Lombok
  - = ; [NHM card]
- glabrius Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 290; TL: Flores
  - = ; [NHM card]
- budinus Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 290; TL: Alor
  - = ; [NHM card]
- Yphthima[sic] aphnius; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4): 292, pl. 17, f. 8-9
- Ypthima leuce; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 28, pl. 1, f. 21-22
- Ypthima aphnius; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 15; [BOW]: pl. 172, f. 13
- Ypthima (Thymipa) aphnius; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Y. a. aphnius Timor, Wetar
  - Ypthima aphnius aphnius; [BAUR], 265 (text)
Ypthima lisandra (Cramer, [1780]) Hong Kong, Tonking, Peninsular Malaya - Indo China, S.China, Taiwan. See [maps]
- Papilio lisandra Cramer, [1780]; Uitl. Kapellen 4 (25-26a): 11, pl. 293, f. G,H; TL: S.China
- Yphthima[sic] micrommatus Holland, 1887; Trans. Amer. Ent. Soc. 14: 115, pl. 2, f. 3
  - = ; [NHM card]
- Ypthima lisandra lisandra, Butterflies in Indo-China [Yutaka Inayoshi]
- Ypthima micrommatus; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 49
- Ypthima lisandra; [BOW]: pl. 205, f. 5; [BOR], 472 (note); [MRS], 392
- Ypthima (Thymipa) lisandra; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Y. l. lisandra N.Vietnam, S.China
- Y. l. bara Evans, 1923 Sikkim - Burma, Thailand, Laos
  - Ypthima avanta bara Evans, 1923; J. Bombay nat. Hist. Soc. 29 (3): 786, 797, No. D. 14/12b
  - Ypthima lisandra bara, Butterflies in Indo-China [Yutaka Inayoshi]
Ypthima singala Felder, 1868 Ceylon. See [maps]
- Ypthima singala Felder, 1868; Verh. zool.-bot. Ges. Wien 18 (1-2): 283; TL: Bholahat, Ceylon, Kumaon
- Ypthima singala; Moore, [1880], Lepid. Ceylon 1 (1): 24, pl. 12, f. 3, 3a; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 31, pl. 1, f. 26, 26a; [BOR], 466
- Ypthima (Thymipa) singala; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima avanta Moore, [1875] Ceylon, Peninsular India, Himalayas - Kashmir, Assam. See [maps] Jewel Fourring
- Yphthima[sic] avanta Moore, [1875]; Proc. zool. Soc. Lond. 1874 (4): 567; TL: Bagh Valley, Cashmere
- Ypthima thora Moore, [1880]; Lepid. Ceylon 1 (1): 24, pl. 12, f. 4, 4a; TL: Ceylon
  - = ?Ypthima singala ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 32
  - = ; [NHM card]
- Ypthima ordinata Butler, 1880; Proc. zool. Soc. Lond. 1880: 148, pl. 15, f. 3
  - = ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 33
- Ypthima avanta; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 33, pl. 1, f. 27; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 472; [BOW]: pl. 172, f. 10; [BOR], 466
- Ypthima (Thymipa) avanta; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Subgenus Ypthima Hübner, 1818

Ypthima Hübner, 1818; Zuträge Samml. exot. Schmett. 1: 17 [missid. TS Papilio philomela Linnaeus]; TS: Ypthima huebneri Kirby

Ypthima; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

The arctous species-group

Arctous; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Arctoa; Shirôzu & Shima, 1979, (incl.)
Serasa; Shirôzu & Shima, 1979, (incl.)

Ypthima arctous (Fabricius, 1775) Australia. See [maps] Dingy Ring
- Papilio arctous Fabricius, 1775; Syst. Ent.: 489
- Yphthima[sic] arctoides Hewitson, 1865; Trans. ent. Soc. Lond. (3) 2 (4): 284
- Ypthima arctous, Lepidoptera Larvae of Australia [Don Herbison-Evans]
- Yphthima[sic] arctous; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4): 283
- Ypthima arctoa;
- Ypthima arctous; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 14, pl. 3, f. 49
- Ypthima (Ypthima) arctous; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Misidentified/misapplied:
- Papilio acrctous; Donovan, 1805, Epitome nat. Hist. Ins. New Holland: pl. 24
Larva on Digitaria didactyla, Imperata cylindrica, I. sp., Themeda triandra, (Unidentified grasses) K. L. & L. E. Dunn, 1991, Review Austr. Butts. (1-4): (120-140)
- Y. a. arctous Northern Territory, Torres Straits Is., Cape York - S.New South Wales
  - Xois arctous arctous
  - Xois arctous arctous; [BAUR], 265
- Y. a. subarctua Tryon, 1890 Papua, New Guinea
  - papuana Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9: 286
  - Xois arctous papuana; [BAUR], 265
Ypthima sesara (Hewitson, 1865) Fiji
- Xois sesara Hewitson, 1865; Trans. ent. Soc. Lond. (3) 2 (4): 282, pl. 17, f. 3-4; TL: Fiji
- Acrophthalmia ? diophthalma Prittwitz, 1867; Stettin ent. Ztg 28 (7-9): 274
  - = ; [NHM card]
- Xois sesara; Fruhstorfer, 1902, Stettin ent. Ztg 63 (1): 354; [BOW]: pl. 172, f. 4; [BAUR], 265
- Ypthima (Ypthima) sesara; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima fulvida (Butler, 1883) Fiji
- Xois fulvida Butler, 1883; Ann. Mag. nat. Hist. (5) 11 (66): 411; TL: Viti-Levu
- vitiensis Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 294
  - = ; [NHM card]
- Xois fulvida; Fruhstorfer, 1902, Stettin ent. Ztg 63 (1): 353; [BAUR], 265
- Ypthima (Ypthima) fulvida; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

The nareda species-group

Nareda; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima nareda (Kollar, [1844]) Pakistan, NW.India, Nepal, Bhutan, Sikkim. See [maps] Large Three-Ring
- Satyrus nareda Kollar, [1844]; in Hügel, Kaschmir und das Reich der Siek 4: 451
- Ypthima nareda; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 20, pl. 1, f. 2, pl. 2, f. 40; [BOW]: pl. 173, f. 4; [BOR], 470; [MRS], 394
- Ypthima (Ypthima) nareda; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Ypthima nareda; Huang & Wu, 2003, Neue Ent. Nachr. 55: 122 (note)
Misidentified/misapplied:
- Ypthima newara; Lee, 1982, in Huang, Insects of Xizang, 2: 135, No. 23 (65) part (missid.)
Larva on Gramineae [MRS]
- Y. n. nareda W.Himalayas
  - Ypthima nareda nareda; [BOR], 470
- Y. n. sarcaposa Fruhstorfer, 1911² Burma, Thailand, Indo China
  - nareda sarcaposa Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 291; TL: Burma
  - Ypthima newara sarcaposa; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 477, pl. 32-33, f. 4
  - Ypthima nareda sarcaposa; [BOR], 470; [MRS], 394
- Y. n. fusca Elwes & Edwards² Assam
  
  = Ypthima fusca Elwes & Edwards, 1893?

The asterope species-group

Asterope; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima asterope (Klug, 1832)

Lebanon, Syria, Himalayas, Kashmir, Baluchistan, India, Tropical Africa. See [maps] Common Three-Ring

Hipparchia asterope Klug, 1832; in Ehrenberg, Symbolae phys., Ins. 3: pl. 29, f. 11-14
Ypthima alemola Swinhoe, 1885; Proc. Zool. Soc. Lond. 1885: 127
- = ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 12
Ypthima complexiva Swinhoe, 1886; Proc. zool. Soc. Lond. 1886 (4): 423, pl. 40, f. 2
- = ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 12
hereroica Grünberg, 1910²; Denkschr. med.-naturw. Ges. Jena 16 (4): 101
- = ; [NHM card]
merlenii Kattulas & Koutsaftikis, 1976; Biologia gallo-hellen 7 (1-2): 156; TL: Greece
- = ; [NHM card]

Ypthima asterope, Lepiforum [LEPIFORUM]
Ypthima asterope, Moths and Butterflies of Europe [MBE]

Yphthima[sic] asterope; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4): 283
Ypthima asterope ab. simpliciocellata; Strand, 1909, Ent. Zs. 23: 112
Ypthima asterope; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 11, pl. 1, f. 4, pl. 3, f. 44b; Strand, 1909, Ent. Zs. 23: 112; Kielland, 1982, Tijdschr. Ent. 125 (5): 104; [BOW]: pl. 204, f. 18; [BOR], 464; [AfrL]; [Otakar Kudrna]
Ypthima (Ypthima) asterope; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

902x657 (~199Kb) underside Ethiopia, Oromia, Debre Libanos env., at German Ethio Lodge, the Gur River right bank. 30th July 2012, Photo © Oleg Kosterin

880x615 (~228Kb) underside Ethiopia, Oromia, Debre Libanos env., at German Ethio Lodge, the Gur River right bank. 30th July 2012, Photo © Oleg Kosterin

1100x1100 (~375Kb) upperside Ethiopia, Oromia, Debre Libanos env., at German Ethio Lodge, the Gur River right bank. 30th July 2012, Photo © Oleg Kosterin

Larva on Cynodon [BK]

Y. a. asterope India, Arabia, N.Tropical Africa
- Ypthima asterope ab. binucholata Strand, 1909; Ent. Zs. 23: 112; TL: Sadani, Tanzania
  - = ?Ypthima granulosa ; Kielland, 1982, Tijdschr. Ent. 125 (5): 104
- Ypthima asterope ab. biocelligera Strand, 1909; Ent. Zs. 23: 112; TL: Sadani, Tanzania
  - = ?Ypthima granulosa ; Kielland, 1982, Tijdschr. Ent. 125 (5): 104
- Ypthima asterope asterope; Kielland, 1982, Tijdschr. Ent. 125 (5): 104, pl. 1, f. 1-8; [BK]: 274, pl. 30, f. 428; [BAFR], 192; [AfrL]
Y. a. mahratta Moore, 1884 NW.India - Assam
- Ypthima mahratta Moore, 1884; J. asiat. Soc. Bengal 53 Pt.II (1): 16; TL: Deccan
  - = Ypthima asterope ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 12
- Ypthima asterope mahratta; [BOR], 464
Y. a. hereroica van Son, 1955 SW.Africa, Nambibia
- Ypthima asterope hereroica van Son, 1955; Transv. Mus. Mem. 8: 156, f. 165
- Yphtima asterope ab. hereroica Grünberg, 1910; Denkschr. med.-naturw. Ges. Jena 16 (4): 101; TL: Damaraland
- Ypthima asterope hereroica; Kielland, 1982, Tijdschr. Ent. 125 (5): 107, pl. 2, f. 1-2; [BAFR], 192 (text); [AfrL]

Ypthima norma (Westwood, 1851) -. See [maps]
- Iphthima norma Westwood, 1851; Gen. diurn. Lep. (2): pl. 67, f. 1
- Ypthima norma; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 51
- Ypthima (Ypthima) norma; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 183
Larva on Bambusa , Cynodon , Digitaria , Ehrharta , Imperata , Microstegium , Oplismenus , Paspalum , Pogonatherum Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 183
- Y. n. burmana Evans, 1923 Burma, Thailand, Laos
  - Ypthima asterope burmana Evans, 1923; J. Bombay nat. Hist. Soc. 29 (3): 785; TL: N.Burma
  - Ypthima norma burmana, Butterflies in Indo-China [Yutaka Inayoshi]
  - Ypthima asterope burmana; [BOR], 464
- Y. n. annamitica Fruhstorfer, 1911 S.Vietnam
  - Ypthima asterope annamitica Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9: 286; TL: Annam
  - Ypthima norma annamitica, Butterflies in Indo-China [Yutaka Inayoshi]
  - Ypthima asterope annamitica; [BOR], 464
- Y. n. pusilla Fruhstorfer, 1911² Sulawesi, Buton, Buru, Ambon
  - norma pusilla Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 287
  - Ypthima pusilla; [BAUR], 265 (text)
  - Ypthima (Ypthima) norma pusilla; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184
Ypthima cataractae van Son, 1955 Victoria Falls, Zambesi
- Ypthima cataractae van Son, 1955; Transv. Mus. Mem. 8: 154 [wet-season]
- Ypthima cataractae f. irvingi van Son, 1955; Transv. Mus. Mem. 8: 154 [dry season]
Ypthima yatta Kielland, 1982 Ethiopia, Kenya. See [maps] Yatta Three-Ring
- Ypthima yatta Kielland, 1982; Tijdschr. Ent. 125 (5): 108, pl. 2, f. 3-6; TL: Kenya, Thika, Yatta
- Ypthima yatta; [BK]: 275; [AfrL]
Ypthima congoana Overlaet, 1955 Zaire (Katanga). See [maps]
- Ypthima asterope congoana Overlaet, 1955; Miss. de Witte 27: 85, f. 12a-b; TL: Upemba, Katanga
- Ypthima congoana; Kielland, 1982, Tijdschr. Ent. 125 (5): 109, pl. 8, f. 1-2, 4-5; [AfrL]
Ypthima rhodesiana Carcasson, 1961 Zambia, Zimbabwe, S.Zaire, Tanzania, SE.Kenya. See [maps]
- Ypthima rhodesiana Carcasson, 1961; Occ. Pap. Coryndon meml Mus. (7): 14, f. 11; TL: Choma, Zambia
- Ypthima rhodesiana; Kielland, 1982, Tijdschr. Ent. 125 (5): 111, pl. 4, f. 1-6; [BK]: 275, pl. 30, f. 430; [AfrL]
Ypthima simplicia Butler, 1876 Ethiopia, S.Sudan, Kenya, N.Tanzania. See [maps]
- Ypthima simplicia Butler, 1876; Ann. Mag. nat. Hist. (4) 18 (108): 481; TL: Ethiopia
- Ypthima simplicia; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 13, pl. 3, f. 54; Kielland, 1982, Tijdschr. Ent. 125 (5): 113, pl. 2, f. 7-8, pl. 3, f. 1-6; [BK]: 275; [AfrL]
Ypthima condamini Kielland, 1982 -. See [maps]
- Ypthima condamini Kielland, 1982; Tijdschr. Ent. 125 (5): 115; TL: Tanzania, MPanda, Sibweza
- Ypthima condamini condamini; [BK]: 276
- Ypthima condamini; [AfrL]
- Y. c. condamini Ethiopia, S.Sudan, E.Africa, Zambia, Mozambique, Transvaal, W.Zaire, N.Angola
  - Ypthima condamini condamini; Kielland, 1982, Tijdschr. Ent. 125 (5): 115, pl. 4, f. 7-14; [AfrL]
- Y. c. nigeriae Kielland, 1982 Nigeria - Ghana, Upper Volta, Senegal
  - Ypthima condamini nigeriae Kielland, 1982; Tijdschr. Ent. 125 (5): 116, pl. 4, f. 15-18; TL: Nigeria, Farniso near Kano
  - Ypthima condamini nigeriae; [AfrL]
Ypthima antennata van Son, 1955 E.Africa, S.Africa, W.Kenya, S.Sudan, Nigeria, Ghana. See [maps]
- Ypthima antennata van Son, 1955; Transv. Mus. Mem. 8: 156, pl. 5-6, f. 86-87; TL: Munnik, N. Transvaal
- Ypthima antennata; Kielland, 1982, Tijdschr. Ent. 125 (5): 118; [AfrL]
- Y. a. antennata E.Africa - S.Africa, Zambia, Uganda, S.Sudan
  - Ypthima antennata antennata; Kielland, 1982, Tijdschr. Ent. 125 (5): 118, pl. 5, f. 1-4; [BK]: 276, pl. 30, f. 433; [AfrL]
- Y. a. cornesi Kielland, 1982 Nigeria, Ghana
  - Ypthima antennata cornesi Kielland, 1982; Tijdschr. Ent. 125 (5): 120; TL: N. Nigeria, Benue Plateau, Jos
  - Ypthima antennata cornesi; [AfrL]
Ypthima jacksoni Kielland, 1982 Kenya (coast) - Ethiopia. See [maps]
- Ypthima jacksoni Kielland, 1982; Tijdschr. Ent. 125 (5): 120, pl. 3, f. 5-8; TL: Kenya, Mombasa, Mtwapa Creek
- Ypthima jacksoni; [BK]: 276; [AfrL]
Ypthima vuattouxi Kielland, 1982 Ivory Coast, Ghana, Nigeria, Cameroun. See [maps]
- Ypthima vuattouxi Kielland, 1982; Tijdschr. Ent. 125 (5): 121, pl. 5, f. 5-8; TL: Ivory Coast, Lamto (Toumodi)
- Ypthima vuattouxi; [AfrL]
Ypthima lamto Kielland, 1982 Ivory Coast, Cameroun. See [maps]
- Ypthima lamto Kielland, 1982; Tijdschr. Ent. 125 (5): 123, pl. 5, f. 9-14; TL: Ivory Coast, Lamto (Toumodi)
- Ypthima lamto; [AfrL]
Ypthima recta Overlaet, 1955 W.Africa - W.Kenya, Zaire, Angola. See [maps]
- Ypthima recta Overlaet, 1955; Miss. de Witte 27: 87, f. 25a-b; TL: Upemba, Zaire
- Ypthima recta; Kielland, 1982, Tijdschr. Ent. 125 (5): 124, pl. 8, f. 3, 6; [BK]: 276; [AfrL]
Ypthima granulosa Butler, 1883 Moçambique, Malawi, Zambia, Rhodesia, N.Botswana, E.Africa. See [maps]
- Ypthima granulosa Butler, 1883; Ann. Mag. nat. Hist. (5) 12 (68): 101; TL: Victoria Nyanza
- Ypthima cataracta van Son, 1955; Transv. Mus. Mem. 8: 154, pl. 5-6, f. 81
  - = ; Kielland, 1982, Tijdschr. Ent. 125 (5): 126
- Ypthima asterope ab. inocellata Strand, 1909; Ent. Zs. 23: 112; TL: Uluna-Langenburg
  - = ; Kielland, 1982, Tijdschr. Ent. 125 (5): 126
- Ypthima asterope ab. triocellata Strand, 1909; Ent. Zs. 23: 112; TL: Langenburg, N.Nyassa-see
  - = ; Kielland, 1982, Tijdschr. Ent. 125 (5): 126
- Ypthima asterope ab. biocellata Strand, 1909; Ent. Zs. 23: 112; TL: Dar es Salaam, Tanzania
  - = ; Kielland, 1982, Tijdschr. Ent. 125 (5): 126
- Ypthima asterope ab. interrupta Strand, 1909; Ent. Zs. 23: 113; TL: Dar es Salaam
  - = ; Kielland, 1982, Tijdschr. Ent. 125 (5): 126
- Ypthima asterope ab. uniocellata Strand, 1909; Ent. Zs. 23: 113; TL: Dar es Salaam
  - = ; Kielland, 1982, Tijdschr. Ent. 125 (5): 126
- Ypthima granulosa; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 12, pl. 3, f. 44, 44a; Kielland, 1982, Tijdschr. Ent. 125 (5): 126, pl. 6, f. 1-8; [BAFR], 192; [BK]: 277, pl. 30, f. 336; [AfrL]
Larva on Poaceaea [BK]
Ypthima pupillaris Butler, 1888 -. See [maps]
- Ypthima pupillaris Butler, 1888; Proc. Zool. Soc. Lond. 1888: 59; TL: River Dangu, Ganyese Tambu, Eq. Afr.
- Ypthima pupillaris; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 20; Strand, 1909, Ent. Zs. 23: 113; Kielland, 1982, Tijdschr. Ent. 125 (5): 133; [AfrL]
- Y. p. pupillaris Ivory Coast - Nigeria, W.Zaire, Rhodesia
  - Ypthima gazana van Son, 1955; Transv. Mus. Mem. 8: 151, pl. 5, f. 79-80
    
    = Ypthima pupillaris pupillaris ; Kielland, 1982, Tijdschr. Ent. 125 (5): 133
  - Ypthima pupillaris ab. depupillata Strand, 1909; Ent. Zs. 23: 113; TL: Loma
    
    = Ypthima pupillaris pupillaris ; Kielland, 1982, Tijdschr. Ent. 125 (5): 133
  - Ypthima pupillaris ab. macrocellata Strand, 1913; Archiv Naturg. 79 A (7): 150; TL: Cameroun
    
    = Ypthima pupillaris pupillaris ; Kielland, 1982, Tijdschr. Ent. 125 (5): 133
  - Ypthima pupillaris pupillaris; Kielland, 1982, Tijdschr. Ent. 125 (5): 133, pl. 6, f. 9-12; [AfrL]
- Y. p. obscurata Kielland, 1982 E.Africa
  - Ypthima pupillaris obscurata Kielland, 1982; Tijdschr. Ent. 125 (5): 134, pl. 6, f. 13-18; TL: Tanzania, MPanda, Ngondo, 1500m
  - Ypthima pupillaris obscurata; [BK]: 277; [AfrL]
Ypthima impura Elwes & Edwards, 1893 -. See [maps] Impure Ringlet
- Ypthima impura Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1): 23, pl. 3, f. 48; TL: Angola
- Ypthima impura; [AfrL]
- Y. i. impura Ivory Coast - Nigeria, Cameroun, Zaire, Angola
  - Ypthima impura impura; Kielland, 1982, Tijdschr. Ent. 125 (5): 135, pl. 7, f. 1-2; [BAFR], 192 (text); [AfrL]
- Y. i. paupera Ungemach, 1932 Ethiopia, E.Africa - Transvaal, Natal
  - Ypthima pupillaris paupera Ungemach, 1932; Mém. Soc. Sci. nat. phys. Maroc 32: 51; TL: Ethiopia
  - Ypthima pupillaris paupera f. hiemis Ungemach, 1932; Mém. Soc. Sci. nat. phys. Maroc 32: 51; TL: Ethiopia
    
    = Ypthima impura paupera ; Kielland, 1982, Tijdschr. Ent. 125 (5): 137
  - Ypthima impura f. badhami van Son, 1955; Transv. Mus. Mem. 8: 150, pl. 56, f. 77, 79; TL: Rhodesia
    
    = Ypthima impura paupera ; Kielland, 1982, Tijdschr. Ent. 125 (5): 137
  - Ypthima impura paupera; Kielland, 1982, Tijdschr. Ent. 125 (5): 137, pl. 7, f. 3-8; [BAFR], 192 (text); [BK]: 277; [AfrL]
Ypthima pulchra Overlaet, 1954 S.Zaire (Shaba), Tanzania - Uganda. See [maps]
- Ypthima pulchra Overlaet, 1954; Lambillionea 54: 41, f. 1a-b; TL: Katanga, Zaire
- Ypthima pulchra; Kielland, 1982, Tijdschr. Ent. 125 (5): 138, pl. 8, f. 7-8, 10-11; [BAFR], 192 (text); [AfrL]
Ypthima praestans Overlaet, 1954 S.Zaire (Shaba), Angola, W.Zambia. See [maps]
- Ypthima praestans Overlaet, 1954; Lambillionea 54: 43, f. 2; TL: Sankuru
- Ypthima praestans; Kielland, 1982, Tijdschr. Ent. 125 (5): 139, pl. 7, f. 9-12, pl. 8, f. 12; [BAFR], 192; [AfrL]
Ypthima doleta Kirby, 1880 Sierra Leone - Ivory Coast, Ghana, Togo, Nigeria - Zaire, Angola, Uganda, NW.Tanzania. See [maps]
- Ypthima doleta Kirby, 1880; Proc. R. Dublin Soc. (2) 2: 335; TL: Sierra Leone
- Ypthima doleta ab. tripunctata Strand, 1909; Ent. Zs. 23: 114; TL: Cameroun
  - = ; Kielland, 1982, Tijdschr. Ent. 125 (5): 141
- Ypthima doleta ab. quadripunctata Strand, 1909; Ent. Zs. 23: 114; TL: Cameroun
  - = ; Kielland, 1982, Tijdschr. Ent. 125 (5): 141
- Ypthima doleta ab. quinquepunctata Strand, 1909; Ent. Zs. 23: 114; TL: Cameroun
  - = ; Kielland, 1982, Tijdschr. Ent. 125 (5): 141
- Ypthima doleta ab. septemocellata Strand, 1909; Ent. Zs. 23: 114; TL: Cameroun
  - = ; Kielland, 1982, Tijdschr. Ent. 125 (5): 141
- Ypthima doleta ab. strigata Strand, 1913; Archiv Naturg. 79 A (7): 150
  - = ; Kielland, 1982, Tijdschr. Ent. 125 (5): 141
- Ypthima doleta ab. defecta Strand, 1913; Archiv Naturg. 79 A (7): 150; TL: Uelleburg
  - = ; Kielland, 1982, Tijdschr. Ent. 125 (5): 141
- ?Ypthima doleta ab. asteropina Strand, 1913; Archiv Naturg. 79 A (7): 150; TL: Alen
  - = ?; Kielland, 1982, Tijdschr. Ent. 125 (5): 141
- Ypthima doleta ab. dschangensis Strand, 1914; Arch. Naturgesch. 80 A (2): 153; TL: Cameroun
  - = ; Kielland, 1982, Tijdschr. Ent. 125 (5): 141
- Ypthima doleta; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 24, pl. 3, f. 51; Strand, 1909, Ent. Zs. 23: 112; Kielland, 1982, Tijdschr. Ent. 125 (5): 141, pl. 7, f. 13-16; [BAFR], 192; [BOW]: pl. 118, f. 23; [AfrL]
Ypthima diplommata Overlaet, 1954 Katanga, W.Zambia, Angola. See [maps]
- Ypthima diplommata Overlaet, 1954; Lambillionea 54: 44, f. 3; TL: Kafakumba, Katanga
- Ypthima diplommata; Kielland, 1982, Tijdschr. Ent. 125 (5): 142, pl. 7, f. 17-18, pl. 8, f. 13-14; [AfrL]
Ypthima albida Butler, 1888 Nigeria, Cameroons. See [maps] Silver Ringlet
- Ypthima albida Butler, 1888; Proc. Zool. Soc. Lond. 1888: 59; TL: Fóda, Equatorial Africa
- Ypthima albida; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 48; Strand, 1909, Ent. Zs. 23: 113; Kielland, 1982, Tijdschr. Ent. 125 (5): 143; [BOW]: pl. 118, f. 6; [BK]: 277, pl. 30, f. 439; [AfrL]
- Y. a. albida W.Kenya, Uganda, NW.Tanzania, E.Zaire, Rwanda, Burundi
  - Ypthima albida argentata Bartel, 1905; Novit. Zool. 12: 134; TL: Niarugungu, Ruanda
    
    = Ypthima albida albida ; Kielland, 1982, Tijdschr. Ent. 125 (5): 143
  - Ypthima albida ab. conradsi Strand, 1909; Ent. Zs. 23: 113; TL: Ukerewe I., L. Victoria
    
    = Ypthima albida albida ; Kielland, 1982, Tijdschr. Ent. 125 (5): 143
  - Ypthima albida argentata; [BAFR], 193 (text)
  - Ypthima albida albida; Kielland, 1982, Tijdschr. Ent. 125 (5): 143; [BAFR], 193; [AfrL]
- Y. a. uniformis Bartel, 1905 E.Zaire (Kivu)
  - Ypthima albida uniformis Bartel, 1905; Novit. Zool. 12: 134; TL: Central Africa
  - Ypthima albida uniformis ab. pseudalbida Dufrane, 1945; 94; TL: Kamitunga
    
    = Ypthima albida uniformis ; Kielland, 1982, Tijdschr. Ent. 125 (5): 144
  - Ypthima albida uniformis ab. impunctata Dufrane, 1945; 94; TL: Kamitunga
    
    = Ypthima albida uniformis ; Kielland, 1982, Tijdschr. Ent. 125 (5): 144
  - Ypthima albida uniformis; Kielland, 1982, Tijdschr. Ent. 125 (5): 144; [BAFR], 193; [AfrL]
- Y. a. occidentalis Bartel, 1905 Camoroun - N.Nigeria
  - Ypthima albida occidentalis Bartel, 1905; Novit. Zool. 12: 134; TL: N. Cameroun
  - Ypthima albida ab. argentoides Strand, 1914; Arch. Naturgesch. 80 A (2): 153; TL: Dschang, Cameroun
    
    = Ypthima albida occidentalis ; Kielland, 1982, Tijdschr. Ent. 125 (5): 143
  - Ypthima albida occidentalis; Kielland, 1982, Tijdschr. Ent. 125 (5): 143; [BAFR], 193 (text); [AfrL]

The chenu species-group

Chenu; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Praenubila; Shirôzu & Shima

Ypthima chenu (Guérin-Méneville, 1843) Coorg, S.India. See [maps] Nilgiri Fourring
- Satyrus chenu Guérin-Méneville, 1843; in Delessert, Souvenirs Voy. Inde (2): 77, pl. 21, f. 2
- Yphthima[sic] chenu; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4): 285
- Ypthima chenui[sic, recte chenu]; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 43; [BIR]; [BOW]: pl. 172, f. 14
- Ypthima chenu; [BOR], 466
- Ypthima (Ypthima) chenu; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima ypthimoides (Moore, 1881) S.India - S.Nilgiris. See [maps] Palni Fourring
- Callerebia ypthimoides Moore, 1881; Trans. ent. Soc. Lond. 1881 (3): 307
- Ypthima robinsoni Distant, 1882; Ann. Mag. nat. Hist. (5) 10 (59): 406
  - = ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 43
- Ypthima ypthimoides; [BOR], 468
- Ypthima (Ypthima) ypthimoides; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima praenubila Leech, 1891 W.China, C.China. See [maps]
- Ypthima praenubila Leech, 1891; Entomologist 24 (Suppl.): 66; TL: ;Ta-chien-lu, Chia-Kow-Ho, Moupin; Omei-Shan; Wa-Shan; Kiukiang
- Ypthima praenubila praenubila, Butterflies in Indo-China [Yutaka Inayoshi]
- Ypthima praenubila; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 18, pl. 3, f. 47; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 481; [BOW]: pl. 205, f. 2; [BOR], 468; [MRS], 393
- Ypthima (Ypthima) praenubila; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Larva on Pogonatherum crinitinum [MRS]
- Y. p. kanonis Matsumura, 1929 Taiwan
  - Ypthima praenubila kanonis Matsumura, 1929; Insecta Matsumurana 3 (4): 141; TL: Formosa
  - Ypthima praenubila kanonis; Uemura, 1984, Tyô to Ga 35 (3): 136; [MRS], 393

The huebneri species-group

Huebneri; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima huebneri Kirby, 1871 NW.India - Peninsular Malaya, Langkawi Is., Singapore, Thailand, S.Yunnan. See [maps] Common Fourring
- Ypthima hübneri [= huebneri] Kirby, 1871; Syn. Cat. diurn. Lep.: 95 (repl. Ypthima philomela Hübner, [1813])
- Ypthima philomela Hübner, [1813]; Zuträge Samml. exot. Schmett. 1: pl. [15], f. 83-84
- Ypthima howra Moore, 1884; J. asiat. Soc. Bengal 53 Pt.II (1): 17; TL: Calcutta
  - = ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 9
- Ypthima apicalis Moore, 1884; J. asiat. Soc. Bengal 53 Pt.II (1): 17; TL: "Deyra Dhoon"
  - = ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 9
- Ypthima kasmira Moore, 1884; J. asiat. Soc. Bengal 53 Pt.II (1): 17; TL: Cashmere
- Ypthima catharina Butler, 1886; Ann. Mag. nat. Hist. (5) 18 (105): 183; TL: Upper Burma
  - = ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 9
- Ypthima jocularia Swinhoe, 1889; Proc. zool. Soc. Lond. 1889 (4): 396; TL: Mahabalesvar
- honora Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 287
  - = ; [NHM card]
- Ypthima huebneri, Lepiforum [LEPIFORUM]
- Ypthima huebneri, Butterflies in Indo-China [Yutaka Inayoshi]
- Yphthima[sic] philomela; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4): 284
- Ypthima hübneri; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 9, pl. 1, f. 5
- Ypthima kasmira; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 10, pl. 1, f. 6
- Ypthima (Ypthima) kasmira; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Ypthima jocularia; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 50
- Ypthima ceylonica huebneri; [BOR], 464
- Ypthima huebneri; [EBW]; [BOW]: pl. 173, f. 2; [BMP]: 135, pl. 17, f. 30-31
- Ypthima (Ypthima) huebneri; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
769x513 (~84Kb) underside Thailand, Krabi 24.12.2003-7.1.2004, Photo © Olli Vesikko
Ypthima ceylonica Hewitson, 1865 Ceylon. See [maps] White Fourring
- Yphthima[sic] ceylonica Hewitson, 1865; Trans. ent. Soc. Lond. (3) 2 (4): 288, pl. 18, f. 14-15; TL: Ceylon
- gellia Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 287, pl. 99 e
  - = ; [NHM card]
- Ypthima ceylonica, Lepiforum [LEPIFORUM]
- Ypthima ceylonica; Moore, [1880], Lepid. Ceylon 1 (1): 25, pl. 12, f. 5, 5a; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 11, pl. 1, f. 8; [BOW]: pl. 172, f. 12
- Ypthima ceylonica ceylonica; [BOR], 464
- Ypthima (Ypthima) ceylonica; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

The megalomma species-group

Megalomma; Shirozu & Shima, 1979; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80; Huang & Wu, 2003, Neue Ent. Nachr. 55: 121
Akbar; Aoki & Uémura, 1984

Ypthima chinensis Leech, 1892 C.China. See [maps]
- Ypthima newara var. chinensis Leech, [1892]; Butts China Japan Corea (1): 89, pl. 10, f. 5
- Ypthima (Ypthima) chinensis; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Ypthima chinensis; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 477; [MRS], 394; Huang & Wu, 2003, Neue Ent. Nachr. 55: 120 (note)
Ypthima iris Leech, 1891 -. See [maps]
- Ypthima iris Leech, 1891; Entomologist 24 (Suppl.): 57; TL: Wau-ssu-kou; Chow-pin-sa
- Ypthima dromonides Oberthür, 1891; Étud. d'Ent. 15: 15, pl. 2, f. 14
  - = ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 42
- hygrophilus Mell, 1942; Arch. Naturgesch. (N.F.) 11: 261
  - = ; [NHM card]
- Ypthima iris hygrophilus; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 485
- Ypthima iris; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 42; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 485; [BOW]: pl. 205, f. 3
- Ytphima iris; [MRS], 395
- Ypthima (Ypthima) iris; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Y. i. iris Sichuan
  - Ypthima iris iris; Huang, 2003, Neue Ent. Nachr. 55: 53 (note)
- Y. i. microiris Uémura & Koiwaya, 2000 NE.Tibet
  - Ypthima iris microiris Uémura & Koiwaya, 2000; ; TL: Qamdo, NE.Tibet
- Y. i. paradromon Uémura & Koiwaya, 2000 N.Yunnan
  - Ypthima iris paradromon Uémura & Koiwaya, 2000; ; TL: N.Yunnan
- Y. i. naqialoa Huang, 2003 Yunnan
  - Ypthima iris naqialoa Huang, 2003; Neue Ent. Nachr. 55: 53, pl. 7, f. 4; TL: on path between Naqialo and Longpo, Nujiang valley, NW.Yunnan
Ypthima dromon Oberthür, 1891 Yunnan. See [maps]
- Ypthima dromon Oberthür, 1891; Étud. d'Ent. 15: 15, pl. 2, f. 12; TL: Yunnan
- Ypthima dromon; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 51; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 488, pl. 32-33, f. 5-6; [MRS], 395
- Ypthima (Ypthima) dromon; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima megalomma Butler, 1874 -. See [maps]
- Ypthima megalomma Butler, 1874; Cistula ent. 1 (9): 236
- Ypthima megalomma; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 44; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 491; [BOW]: pl. 204, f. 4 (text only); [MRS], 395
- Ypthima (Ypthima) megalomma; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Y. m. megalia de Nicéville, 1897 N.Shan States
  - Ypthima megalia de Nicéville, 1897; J. Asiat. Soc. Bengal 66 Pt.II (3): 546, pl. 1, f. 5
  - Ypthima megalia; [BOR], 472 (note)
Ypthima beautei Oberthür, 1884 -. See [maps]
- Ypthima beautei Oberthür, 1884; Étud. d'Ent. 9: 18, pl. 2, f. 1; TL: Ta-tsien-lo, West China
- Ypthima (Ypthima) beautei; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Ypthima beautei; [BOW]: pl. 205, f. 4 (text only); Huang & Wu, 2003, Neue Ent. Nachr. 55: 115 (note)
- Y. b. beautei W.China
  - Ypthima beautei beautei; Huang & Wu, 2003, Neue Ent. Nachr. 55: 118 (note), f. 8, 9, 11, pl. 10, f. 1
- Y. b. qinghaiensis Huang & Wu, 2003 Qinghai
  - Ypthima beautei qinghaiensis Huang & Wu, 2003; Neue Ent. Nachr. 55: 118, f. 5-7, 12, pl. 10, f. 2; TL: Make valley, E.Qinghai, China
Ypthima pseudodromon Forster, [1948] NW.Yunnan. See [maps]
- Ypthima pseudodromon Forster, [1948]; Mitt. Münch. Ent. Ges. 34 (2): 488, pl. 32-33, f. 7-8; TL: Likiang, N.Yunnan
- Ypthima (Ypthima) pseudodromon; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Ypthima pseudodromon; Huang & Wu, 2003, Neue Ent. Nachr. 55: 116 (ntoe)
Ypthima akbar Talbot, 1947 Burma, Thailand, Laos, S.Yunnan. See [maps]
- Ypthima akbar Talbot, 1947; Fauna Br. Ind., Butt. (2nd ed.) 2: 324; TL: Karen Hills
- Ypthima akbar, Butterflies in Indo-China [Yutaka Inayoshi]
- Ypthima (Ypthima) akbar; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima insolita Leech, 1891 W.China. See [maps]
- Ypthima insolita Leech, 1891; Entomologist 24 (Suppl.): 66; TL: Wa-ssu-Kou
- Ypthima insolita; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 45; [BOW]: pl. 205, f. 4
- Ypthima (Ypthima) insolita; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima putamdui South, 1913 Sichuan. See [maps]
- Ypthima putamdui South, 1913; J. Bomb. nat. Hist. Soc. 22 (2): 348; TL: Nya Chua, Sichuan
- Ypthima (Ypthima) putamdui; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Ypthima putamdui; Huang & Wu, 2003, Neue Ent. Nachr. 55: 116 (note)
Ypthima baileyi South, 1913 Sichuan. See [maps]
- Ypthima baileyi South, 1913; J. Bomb. nat. Hist. Soc. 22 (2): 348; TL: Batang, W.Sichuan
- Ypthima baileyi; Huang & Wu, 2003, Neue Ent. Nachr. 55: 118, f. 7, 14, 18-20, pl. 10, f. 4-5
Ypthima frontierii Uémura & Monastyrskii, 2000 N.Vietnam, Guanxi. See [maps]
- Ypthima frontierii Uémura & Monastyrskii, 2000; Trans. Lepid. Soc. Japan 51 (2): 150, f. 1a-c, 2, 4-9; TL: N.Vietnam, Lao Cai Prov., Sa Pa, ~2000m (103°46'E, 22°15'N)
- Ypthima frontierii, Butterflies in Indo-China [Yutaka Inayoshi]
- Ypthima frontierii; Huang & Wu, 2003, Neue Ent. Nachr. 55: 120, f. 21, pl. 10, f. 7
Ypthima yoshinobui Huang & Wu, 2003 Qinghai
- Ypthima yoshinobui Huang & Wu, 2003; Neue Ent. Nachr. 55: 115, f. 3, 4, 7, 10, pl. 10, f. 3; TL: Make valley, E.Qinghai, China
Ypthima lihongxingi Huang & Wu, 2003 Hubei. See [maps]
- Ypthima lihongxingi Huang & Wu, 2003; Neue Ent. Nachr. 55: 120, f. 13, 15-17, pl. 10, f. 6; TL: Xing-dou-shan, Lichuan, Hubei

The newara species-group

Newara; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80; Huang, 2001, Neue Ent. Nachr. 51: 92

Ypthima newara Moore, [1875] Nepal. See [maps]
- Yphthima[sic] newara Moore, [1875]; Proc. zool. Soc. Lond. 1874 (4): 567; TL: Nepal
- Ypthima nareda newara; [BOR], 470
- Ypthima (Ypthima) newara; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Ypthima newara; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 21, pl. 2, f. 42; Huang, 2001, Neue Ent. Nachr. 51: 92 (note), f. 81, pl. 9, f. 72-right
Misidentified/misapplied:
- Yphthima nareda; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4): pl. 17, f. 7 (nec. f. 6)
- Y. n. yaluzangbui Huang, 1999 SE.Tibet
  - Ypthima newara yaluzangbui Huang, 1999; ; TL: Metok
  - Ypthima newara yaluzangbui; Huang, 2001, Neue Ent. Nachr. 51: 92 (note), f. 80 (gen); Huang, 2003, Neue Ent. Nachr. 55: 52
Ypthima dengae Huang, 2001 SE.Tibet. See [maps]
- Ypthima dengae Huang, 2001; Neue Ent. Nachr. 51: 92, f. 79, 115, 117, pl. 7. f. 49, 53; TL: Ba-an-tang, Chayu area Tibet
- Ypthima dengae; Huang, 2003, Neue Ent. Nachr. 55: 52
Ypthima confusa Shirôzu & Shima, 1977 Nepal, N.Thailand, Laos, Yunnan. See [maps]
- Ypthima confusa Shirôzu & Shima, 1977; Kontyu 45 (4): 504, f. 1c-d, 1g; TL: Kalapani, Nepal
- Ypthima confusa, Butterflies in Indo-China [Yutaka Inayoshi]
- Ypthima (Ypthima) confusa; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Ypthima confusa; Huang, 2001, Neue Ent. Nachr. 51: 92 (note); Huang, 2003, Neue Ent. Nachr. 55: 49, f. 71, pl. 6, f. 8-10
- Y. c. confusa
  - Ypthima confusa confusa; Huang, 2001, Neue Ent. Nachr. 51: f. 119 (gen.), pl. 9, f. 72-left
Ypthima muotuoensis Huang, 2000 -. See [maps]
- Ypthima confusa muotuoensis Huang, 2000; ; TL: Metok
- Ypthima confusa muotuoensis; Huang, 2001, Neue Ent. Nachr. 51: f. 84 (gen.)
- Ypthima muotuoensis; Huang, 2003, Neue Ent. Nachr. 55: 48 (note)
- Y. m. muotuoensis SE.Tibet
  - Ypthima muotuoensis muotuoensis; Huang, 2003, Neue Ent. Nachr. 55: 49, f. 70, pl. 6, f. 4-7
- Y. m. dulongae Huang, 2003 Yunnan
  - Ypthima muotuoensis dulongae Huang, 2003; Neue Ent. Nachr. 55: 48, 50, f. 67, 69, pl. 6, f. 1-3; TL: Xiandjiudang to Longyuan, Dulong valley, NW.Yunnan
Ypthima pemakoi Huang, 1998 SE.Tibet. See [maps]
- Ypthima pemakoi Huang, 1998; ; TL: Metok
- Ypthima pemakoi; Huang, 2001, Neue Ent. Nachr. 51: 92 (note), f. 85, 116; Huang, 2003, Neue Ent. Nachr. 55: 49, f. 74, pl. 7, f. 1-2
Ypthima sinica Uémura & Koiwaya, 2000 Sichuan, E.China. See [maps]
- Ypthima sinica Uémura & Koiwaya, 2000; ; TL: Qingchengshan, Sichuan
- Ypthima sinica; Huang, 2001, Neue Ent. Nachr. 51: 92 (note); Huang, 2003, Neue Ent. Nachr. 55: 52, f. 75
Ypthima tiani Huang & Liu, 2000 -. See [maps]
- Ypthima tiani; Huang, 2001, Neue Ent. Nachr. 51: 92 (note)
- Y. t. tiani Sichuan
- Y. t. nuae Huang, 2001 Yunnan
  - Ypthima tiani nuae Huang, 2001; Neue Ent. Nachr. 51: 93, f. 82, 114, 120, pl. 7, f. 50, 54; TL: Longpo to Ninadan, Nujiang Valley, SE.Tibet
  - Ypthima tiani nuae; Huang, 2003, Neue Ent. Nachr. 55: 51, f. 68, 72-73, pl. 6, f. 11-18

The pandocus species-group

Pandocus; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima pandocus Moore, [1858] -. See [maps]
- Ypthima pandocus Moore, [1858]; in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1: 235; TL: Java
- Ypthima pandocus var. mentawica Hagen, 1902; Abh. Senckenb. Nat. Ges. 20: 330; TL: Mentawej
  - = ; [NHM card]
- naerius Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 293; TL: Nias
  - = ; [NHM card]
- nacrianus Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 293; TL: Minahassa
  - = ; [NHM card]
- dyna Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 293
  - = ; [NHM card]
- andokides Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 293; TL: Java
  - = ; [NHM card]
- Ypthima pandocus barissanus Martin, 1929; Mitt. Münch. Ent. Ges. 19 (5-9): 125; TL: Battak
  - = ; [NHM card]
- Ypthima pandocus minimus Martin, 1929; Mitt. Münch. Ent. Ges. 19 (5-9): 126; TL: SE.Borneo
  - = ; [NHM card]
- Ypthima pandocus balianus Martin, 1929; Mitt. Münch. Ent. Ges. 19 (5-9): 127; TL: Bali I.
  - = ; [NHM card]
- Ypthima pandocus corticaria ab. kosong Cowan, 1956; Bull. Raffles Mus. 27: 39; TL: Malaya
  - = ; [NHM card]
- microocellata Miller & Miller, 1978; Bull. Allyn Mus. 49: 10; TL: Batjan
  - = ; [NHM card]
- Ypthima pandocus, Lepiforum [LEPIFORUM]
- Yphthima[sic] pandocus; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4): 290, pl. 18, f. 12; van Eecke, 1913, Notes Leyden Mus. 35: 243
- Ypthima pandocus; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 22, pl. 2, f. 38; [BOR], 468
- Ypthima (Ypthima) pandocus; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Y. p. pandocus Java
  - Ypthima pandocus pandocus; [BOR], 468
- Y. p. sertorius Fruhstorfer, 1911² Borneo
  - pandocus sertorius Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 293, pl. 99 e
  - Ypthima pandocus sertorius; [BOR], 468
- Y. p. corticaria Butler, 1879 Sumatra, Peninsular Malaya Common Three-ring
  - Ypthima corticaria Butler, 1879; Trans. linn. Soc. Lond. (2) 1 (8): 537; TL: Malacca
    
    = Ypthima pandocus ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 22
  - Ypthima pandocus emporialis Martin, 1929; Mitt. Münch. Ent. Ges. 19 (5-9): 126; TL: "Bukit Fima", Singapore
  - Ypthima pandocus f. alada Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9: 293; TL: Battaker Plateau, NE.Sumatra.
  - Ypthima pandocus corticaria, Butterflies in Indo-China [Yutaka Inayoshi]
  - Ypthima pandocus corticaria; [BMP]: 135, pl. 17, f. 16-17; [BOR], 468
- Y. p. tahanensis Pendlebury, 1933 Peninsular Malaya (Gunong Tahan)
  - Ypthima tahanensis Pendlebury, 1933; J. Fed. Malay St. Mus. 17 (2): 384
  - Ypthima pandocus tahanensis; Eliot, 1959, Bull. Br. Mus. nat. Hist. (Ent.) 7 (8): 372; [BMP]: 135, pl. 17, f. 18-19; [BOR], 468
- Y. p. moenus Fruhstorfer, 1911² Natuna Is.
  - pandocus moenus Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 293; TL: Natuna Is.
  - Ypthima pandocus moenus; [BOR], 468
- Y. p. aquillius Fruhstorfer, 1911² Palawan
  - pandocus aquillius Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 293; TL: Palawan
  - Ypthima pandocus aquillius; [BOR], 468
- Y. p. calamus Fruhstorfer, 1911² ?Sulu Is.
  - pandocus calamus Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 293
  - Ypthima pandocus calanus[sic?]; [BOR], 468
- Y. p. jamaeus Fruhstorfer, 1911² Banggai
  - pandocus jamaeus Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 293
  - Ypthima pandocus jamaeus; [BOR], 468
- Y. p. chaboras Fruhstorfer, 1911² Philippines (Basilan, Mindanao)
  - pandocus chaboras Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 293; TL: Bazilan
  - Ypthima pandocus chaboras; [BOR], 468
- Y. p. sempera C. & R. Felder, 1863 Philippines (Luzon, ...)
  - Ypthima sempera C. & R. Felder, 1863; Wien. ent. Monats. 7 (4): 125
    
    = Ypthima pandocus ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 22
  - Ypthima (Ypthima) sempera; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
  - Ypthima pandocus sempera; [BOR], 468
Ypthima hanburyi Holloway, 1984 Sarawak (Gunong Mulu)
- Ypthima hanburyi Holloway, 1984; Sarawak Mus. J. (Suppl. II): -
- Ypthima hanburyi; [BOR], 471
- Ypthima (Ypthima) hanburyi; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima nigricans Snellen, 1892 Java. See [maps]
- Ypthima nigricans Snellen, 1892; Tijdschr. Ent. 35 (3/4): 135; TL: W.Java
- phasis Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 294
  - = ; [NHM card]
- Ypthima nigricans; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 52; [BOR], 468
- Ypthima (Ypthima) nigricans; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Ypthima kalelonda Westwood, 1888 -. See [maps]
- Ypthima kalelonda Westwood, 1888; Trans. Ent. Soc. Lond. 1888 (3): 475, pl. 12, f. 4; TL: Kalelonda
  - = Ypthima pandocus ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 22
- Ypthima kaledonda[sic]; [NHM card]
- Ypthima kalelonda; [BOR], 468
- Ypthima (Ypthima) kalelonda; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184
- Y. k. kalelonda N.Sulawesi, Talisei
  - Ypthima kalelonda kalelonda; [BOR], 468
  - Ypthima (Ypthima) kalelonda kalelonda; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184
- Y. k. celebensis Rothschild, 1892 Sulawesi
  - Ypthima celebensis Rothschild, 1892; Dt. ent. Z. Iris 5 (2): 433, pl. 5, f. 3-4; TL: S.Celebes
    
    = Ypthima loryma ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 23
  - Ypthima (Ypthima) kalelonda celebensis; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184
- Y. k. mangolina Uemura, 1982 Sula
  - Ypthima (Ypthima) kalelonda mangolina; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184
Ypthima risompae Uémura, 1982 Sulawesi. See [maps]
- Ypthima (Ypthima) risompae; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184
Larva on Miscanthus , (in captivity) Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184
Ypthima ancus Fruhstorfer, 1911² S.Sulawesi (Bonthain Peak). See [maps]
- ancus Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 293, pl. 99 g; TL: Bonthein
- Ypthima ancus; [BOR], 472 (note)
- Ypthima (Ypthima) ancus; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184
Ypthima gavalisi Martin, 1913 Sulawesi. See [maps]
- Ypthima gavalisi Martin, 1913; Dt. ent. Z. Iris 27 (2): 109, (3):121; TL: Celebes
- ?pandocoides Miller & Miller, 1978; Bull. Allyn Mus. 49: 11; TL: Celebes
- Ypthima gavalisi; [BOR], 466
- ?Ypthima pandocoides; [NHM card]
- Ypthima gavalisi f. pandocoides; [BOR], 467, f.
- Ypthima gawalisi; (missp.)
- Ypthima (Ypthima) gavalisi; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184
Ypthima loryma Hewitson, 1865 Sulawesi. See [maps]
- Yphthima[sic] loryma Hewitson, 1865; Trans. ent. Soc. Lond. (3) 2 (4): 289, pl. 18, f. 16-17; TL: Macassar, Celebes
  - = Ypthima pandocus ; [NHM card]
- Ythima loryma; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 23, pl. 3, f. 50
- Ypthima loryma; [BOR], 466; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Ypthima (Ypthima) loryma; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184
Ypthima sensilis Kashiwai, 1982 Philippines (Mindanao). See [maps]
- Ypthima sensilis Kashiwai, 1982; Tyô to Ga 33 (1,2): 44; TL: Penangudloton, Upper region of Upian River, Davao City Prov., Mindanao
- Ypthima sensilis; [BOR], 468
- Ypthima (Ypthima) sensilis; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

The motschulskyi species-group

Motschulskyi; Shirôzu & Shima; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80; Huang, 2001, Neue Ent. Nachr. 51: 91
Sordida; Shirôzu & Shima
Obscura; Shirôzu & Shima

Ypthima motschulskyi (Bremer & Grey, 1853) E.China, Hong Kong, Amur, Korea, Japan. See [maps]
- Satyrus motschulskyi Bremer & Grey, 1853; Schmett. N. China: 8, pl. 2, f. 2
- Ypthima akragas var. takamukuana Matsumura, 1919; Thous. Ins. Japan. Addit. 3: 526, pl. 37, f. 4, 4a; TL: Formosa
  - = ; [NHM card]
  - = Ypthima perfecta akragas ; Uemura, 1984, Tyô to Ga 35 (3): 136
- formosicola Matsumura, 1929; Ill. comm. Ins. Japan 1: 10
  - = ; [NHM card]
- Ypthima motschulskyi ab. yoshisakai Murayama, 1961; Tyô to Ga 11 (4): 57, f. 5; TL: Mikage-cho, Kobe
- Yphthima motschulskiji[sic]; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4): 290
- Ypthima motschulskyi; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 16, pl. 2, f. 34; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 479; Kudrna, 1974, Atalanta 5: 99; [BRU], 192; [BOW]: pl. 205, f. 6; [MRS], 394; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 44
- Ypthima (Ypthima) motschulskyi; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
668x560 (~77Kb) Korea, 1989, S.W. Cheong col. & det., Photo © Enrique Garcia-Barros

Larva on Carex [BRU], Fukuda et al., 1983
- Y. m. motschulskyi
  - Ypthima obscura Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1): 17, pl. 2, f. 35; TL: Gensan, Korea
    
    = Ypthima motschulskyi motschulskyi ; Dubatolov & Lvovsky, 1997, Trans. lepid. Soc. Japan 48 (4): 194
  - Ypthima elongatum Matsumura, 1929; Insecta Matsumurana 3 (4): 142; TL: Korea
    
    = Ypthima amphithea obscura ; Uemura, 1984, Tyô to Ga 35 (3): 135
    = Ypthima motschulskyi motschulskyi ; Dubatolov & Lvovsky, 1997, Trans. lepid. Soc. Japan 48 (4): 194
  - Ypthima obscura; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 480, pl. 32-33, f. 11-12
  - Ypthima motschulskyi motschulskyi; Dubatolov & Lvovsky, 1997, Trans. lepid. Soc. Japan 48 (4): 194
- Y. m. amphithea Ménétriés, 1859
  - Ypthima amphithea Ménétriés, 1859; in Schrenck, Reise Forschungen Amur-Lande 2 (1): 41, pl. 3, f. 10; TL: Amoor-land, Sula
    
    = Ypthima motschulskyi ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 16
  - Yphthima[sic] amphithea; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4): 289
  - Ypthima (Ypthima) amphithea; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
  - Ypthima motschulskyi amphithea; Dubatolov & Lvovsky, 1997, Trans. lepid. Soc. Japan 48 (4): 194; Huang, 2001, Neue Ent. Nachr. 51: 90 (note); [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 44
Ypthima yangjiahei Huang, 2001 Yunnan. See [maps]
- Ypthima yangjiahei Huang, 2001; Neue Ent. Nachr. 51: 91, f. 78, 113, pl. 6, f. 45; TL: Langpo to Ninadan, Nujiang Valley, SE.Tibet
- Ypthima yangjiahei; Huang, 2003, Neue Ent. Nachr. 55: 100 (note), f. 161, pl. 7, f. 5-6
Ypthima multistriata Butler, 1883 Taiwan. See [maps]
- Ypthima multistriata Butler, 1883; Ann. Mag. nat. Hist. (5) 12 (67): 50; TL: Ichang
- Ypthima arcuata Matsumura, 1919; Thous. Ins. Japan. Addit. 3: 699, pl. 53, f. 17; TL: Formosa, Horisha
  - = ; Uemura, 1984, Tyô to Ga 35 (3): 135; Dubatolov & Lvovsky, 1997, Trans. lepid. Soc. Japan 48 (4): 194
- Ypthima multistriata; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 18, pl. 2, f. 36; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 480; [BOR], 466; [MRS], 395; Dubatolov & Lvovsky, 1997, Trans. lepid. Soc. Japan 48 (4): 195; Huang, 2001, Neue Ent. Nachr. 51: 91 (note); [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 43
980x901 (~186Kb) underside Russia, Khabarovskiy Kray, Khabarovsk southern env., Korfovskiy settlement. 11th July 2014, Photo © Oleg Kosterin

Larva on Miscanthus sinensis, Setaria palmifolia, Isachne globosa, Paspalum conjugatum [MRS]
- Y. m. ganus Fruhstorfer, 1911 N.China, C.China
  - multistriata ganus Fruhstorfer, 1911; in Seitz, Gross-Schmett. Erde 9: 291; TL: Qingdao
  - Ypthima multistriata koreana Dubatolov & Lvovsky, 1997; Trans. lepid. Soc. Japan 48 (4): 195; TL: Korea
    
    = Ypthima multistriata ganus ; Huang, 2001, Neue Ent. Nachr. 51: 91
  - Ypthima multistriata koreana, Lepidoptera collection [SZMN]
  - Ypthima multistriata koreana; [RFEL]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 43
  - Ypthima motschulskyi ganus; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 479
  - Ypthima multistriata ganus; Huang, 2001, Neue Ent. Nachr. 51: 91 (note)
- Y. m. niphonica Murayama, 1969 Japan (Honshu)
  - Ypthima niphonica Murayama, 1969; Tohoku Konchu Kenkyu 4 (2): 18
  - Ypthima motschulskyi niphonica;
  - Ypthima multistriata niphonica; Dubatolov & Lvovsky, 1997, Trans. lepid. Soc. Japan 48 (4): 196
- Y. m. tsushimana Murayama, 1969 Japan
  - Ypthima niphonica tsushimana Murayama, 1969; Tohoku Konchu Kenkyu 4 (2): 20
  - Ypthima multistriata tsushimana; Dubatolov & Lvovsky, 1997, Trans. lepid. Soc. Japan 48 (4): 196
Ypthima perfecta Leech, 1892 -. See [maps]
- Ypthima motschulskyi var. perfecta Leech, [1892]; Butts China Japan Corea (1): 88, pl. 10, f. 7
- Ypthima (Ypthima) perfecta; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Ypthima perfecta; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 19, pl. 2, f. 37; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 477; [MRS], 393; Huang, 2001, Neue Ent. Nachr. 51: 91 (note)
- Y. p. perfecta W.China, C.China
- Y. p. akragas Fruhstorfer, 1911 Taiwan
  - perfecta akragas (Fruhstorfer, 1911); in Seitz, Gross-Schmett. Erde 9: 291
  - Ypthima perfecta akragas; [BOR], 472 (note); [MRS], 393
  Larva on Miscanthus sinensis, M. transmorrisonensis, Oplismenus undulatifolius [MRS]
Ypthima esakii Shirôzu, 1960 Taiwan
- Ypthima esakii Shirôzu, 1960; Butterflies of Formosa in Colour: 128
- Ypthima esakii; [BOR], 472 (note); [MRS], 394
- Ypthima (Ypthima) esakii; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
Larva on Miscanthus sinensis, Setaria palmifolia, Isachne globosa, Oplismenus undulatifolius [MRS]
Ypthima sordida Elwes & Edwards, 1893 China (N.Yunnan, Shensi, ...). See [maps]
- Ypthima sordida Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1): 19, pl. 3, f. 52; TL: Kiukiang
- Ypthima sordida tsinlingi Forster, [1948]; Mitt. Münch. Ent. Ges. 34 (2): 478, pl. 30-31, f. 14-15; TL: Tapaishan, Tsinlingshan, S.Shensi
  - = ; [NHM card]
- Ypthima (Ypthima) sordida; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Ypthima sordida; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 478, pl. 30-31, f. 12-13; Huang, 2001, Neue Ent. Nachr. 51: 91 (note)
Ypthima lycus de Nicéville, 1889 Assam, Bhutan, Sikkim. See [maps] Plain Three-Ring
- Ypthima lycus de Nicéville, 1889; J. Bombay nat. Hist. Soc. 4 (3): 165, pl. A, f. 2
- ?Ypthima lycoides Watson, 1897; J. Bombay nat. Hist. Soc. 10 (4): 648; TL: North Chin Hills, Burma
- Ypthima (Ypthima) lycus; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Ypthima lycus; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 19, pl. 2, f. 39; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 479, pl. 32-33, f. 9-10; [BOR], 470; Huang, 2001, Neue Ent. Nachr. 51: 91 (note)
Ypthima riukiuana Matsumura, 1906 Japan. See [maps]
- Ypthima riukiuana Matsumura, 1906; Annot. zool. japon. 6 (1): 11, pl. 1, f. 2; TL: Okinawa, The Loochoos
- Ypthima riukiuana; Uemura, 1984, Tyô to Ga 35 (3): 136
- Ypthima (Ypthima) riukiuana; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Ypthima riukiuana; Huang, 2001, Neue Ent. Nachr. 51: 91 (note)
Ypthima imitans Elwes & Edwards, 1893 Hainan. See [maps]
- Ypthima imitans Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1): 17, pl. 3, f. 53; TL: Hainan
- Ypthima imitans, Butterflies in Indo-China [Yutaka Inayoshi]
- Ypthima (Ypthima) imitans; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Ypthima imitans; [MRS], 394; Huang, 2001, Neue Ent. Nachr. 51: 91 (note)
Ypthima phania (Oberthür, 1891) Yunnan. See [maps]
- phania (Oberthür, 1891) (Epinephele); Étud. d'Ent. 15: 17, pl. 2, f. 17; TL: Yunnan
- Ypthima phania; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 492, pl. 32-33, f. 13-14; Huang, 2001, Neue Ent. Nachr. 51: 91 (note)
Ypthima watsoni (Moore, [1893]) Assam, Manipur - S.Burma. See [maps] Looped Three-Ring
- Pandima watsoni Moore, [1893]; Lepidoptera Indica 2 (16): 89, (15) pl. 113, f. 4, 4a-b; TL: Burma
- Ypthima watsoni watsoni, Butterflies in Indo-China [Yutaka Inayoshi]
- Ypthima watsoni; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 47, pl. 1, f. 25; [BOR], 470; Huang, 2001, Neue Ent. Nachr. 51: 91 (note)
- Y. w. watsoni Assam - Burma, Thailand
  - Ypthima watsoni var. howarthi Cantlie & Norman, 1959; J. Bombay nat. Hist. Soc. 56 (1): 66; TL: Sebong, Manipur
- Y. w. peninsulae Aoki & Uémura, 1984 Peninsular Thailand
  - Ypthima watsoni peninsulae Aoki & Uémura, 1984; Tokurana 6/7: 87, f. 16i-j, 17, 43, pl. 5, f. 4, 8; TL: Thailand
  - Ypthima watsoni peninsulae, Butterflies in Indo-China [Yutaka Inayoshi]
- Y. w. inouei Shirôzu & Shima, 1977 S.Vietnam
  - Ypthima inouei Shirôzu & Shima, 1977; Kontyu 45 (4): 507, f. 1e-f; TL: Trang Bom, S.Vietnam
  - Ypthima watsoni inouei, Butterflies in Indo-China [Yutaka Inayoshi]
  - Ypthima (Ypthima) inouei; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
  - ?Ypthima watsoni r. inouei; Huang, 2001, Neue Ent. Nachr. 51: 91
Ypthima fusca Elwes & Edwards, 1893
- Ypthima fusca Elwes & Edwards, 1893; Trans. Ent. Soc. Lond. 1893 (1): 21
- Ypthima (Ypthima) fusca; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Ypthima fusca; Huang, 2001, Neue Ent. Nachr. 51: 91 (note)
Ypthima masakii Ito, 1947 Japan. See [maps]
- Ypthima masakii Ito, 1947; Zephyrus 9: 272; TL: Loochoo I.
- Ypthima (Ypthima) masakii; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Ypthima masakii; Huang, 2001, Neue Ent. Nachr. 51: 91 (note)
Ypthima ciris Leech, 1891 W.China. See [maps]
- Ypthima ciris Leech, 1891; Entomologist 24 (Suppl.): 4; TL: Wa-Shan; Chia-Ting-Fu; Ta-Tsien-Lu; Huang-Mu-Chang
- Ypthima clinia Oberthür, 1891; Étud. d'Ent. 15: 16, pl. 2, f. 13
  - = ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 41
- Ypthima civis[sic]; [NHM card]
- Ypthima (Ypthima) ciris; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80
- Ypthima ciris; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 41; Forster, [1948], Mitt. Münch. Ent. Ges. 34 (2): 486; [BOW]: pl. 205, f. 3 (text only); [MRS], 393; Huang, 2001, Neue Ent. Nachr. 51: 91 (note)
- Y. c. clinioides Oberthür, 1891 Yunnan, ...
  - Ypthima clinioides Oberthür, 1891; Étud. d'Ent. 15: 16
  - Ypthima clinioides; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 51
  - Ypthima ciris clinioides; Huang, 2003, Neue Ent. Nachr. 55: 100 (note)

The ? species-group

?; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima bolanica Marshall, 1882 Baluchistan. See [maps] Desert Fourring
- Ypthima bolanica Marshall, 1882; Proc. zool. Soc. Lond. 1882 (4): 759; TL: Bolan Pass
- Ypthima bolanica; [AfrL]
Ypthima inica Hewitson, 1865 Punjab - C.India, Bengal, Assam. See [maps] Lesser Three-Ring
- Yphthima[sic] inica Hewitson, 1865; Trans. ent. Soc. Lond. (3) 2 (4): 284, pl. 17, f. 5; TL: N.India [?Bengal]
- Yphthima[sic] ariaspa Moore, [1875]; Proc. zool. Soc. Lond. 1874 (4): 568; TL: Punjab
- Ypthima rara Butler, 1883; Proc. zool. Soc. Lond. 1883 (2): pl. 24, f. 1
  - = Ypthima ariaspa ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 11
- Ypthima alkibie Swinhoe, 1886; Proc. zool. Soc. Lond. 1886 (4): 422
  - = ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 13
- Ypthima daedalea Swinhoe, 1886; Proc. zool. Soc. Lond. 1886 (4): 423
  - = Ypthima ariaspa ; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 11
- Ypthima ariaspa; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 11
- Ypthima inica; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 13, pl. 1, f. 9; [BOR], 464
Ypthima abnormis Shelford, 1904 Borneo (Sarawak). See [maps]
- Ypthima abnormis Shelford, 1904; J. Straits Asiat. Soc. 41: 98; TL: Kuching, Sarawak
- Ypthima abnormis; [BOR], 472
Ypthima junkoae Uémura, 1999 C.Sulawesi. See [maps]
- Ypthima (Ypthima) junkoae; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 184

Subgenus ?

?; Shima, 1988, Spec. Bull. lepid. Soc. Japan (6): 80

Ypthima argillosa Snellen, 1892 Java. See [maps]
- Ypthima argillosa Snellen, 1892; Tijdschr. Ent. 35 (3/4): 133; TL: Java
- Ypthima argillosa; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 52; [BOR], 472 (note)
Ypthima florensis Snellen, 1891 Flores. See [maps]
- Yphthima[sic] florensis Snellen, 1891; Tijdschr. Ent. 34: 235, pl. 14, f. 3; TL: Flores
- Ypthima florensis; [BOR], 472 (note)
Ypthima angustipennis Takahashi, 2000 Taiwan
- Ypthima angustipennis Takahashi, 2000; Trans. lepid. Soc. Japan 51 (1): 4; TL: Taiwan, Kaohsiung, Tengzhi
Ypthima hyagriva Moore, 1857 N.India, Kashmir - Kumaon, Sikkim. See [maps] Brown Argus
- Yphthima[sic] hyagriva Moore, 1857; in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1: 236; TL: Darjeeling
- Yphthima[sic] hyagriva; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4): pl. 18, f. 11
- Dallachia hyagriva;
- ?Erebia hyagriva; [BOW]: pl. 200, f. 13
- Callerebia hyagriva; [NHM card]
Ypthima albipuncta Lee, 1985 Yunnan. See [maps]
- Ypthima albipuncta Lee, 1985; ; TL: Yi-liang, Yunnan, 1800m
- Ypthima albipuncta; Huang & Wu, 2003, Neue Ent. Nachr. 55: 122, f. 1-2, pl. 10, f. 8-9
Ypthima posticalis Matsumura, 1909 Taiwan. See [maps]
- Ypthima posticalis Matsumura, 1909; Ent. Zs. 23 (19): 92; TL: Formosa, Horisha
- Ypthima minuta Matsumura, 1909; Ent. Zs. 23 (19): 92; TL: Formosa, Hokulo
  - = ; Uemura, 1984, Tyô to Ga 35 (3): 136
  - = Ypthima lisandra ; [NHM card]
- Ypthima norma posticalis; Uemura, 1984, Tyô to Ga 35 (3): 136
- Ypthima posticalis; [BOR], 472 (note)
- Y. p. aei Shirôzu & Shima, 1977 Philippines (Luzon)
  - Ypthima posticalis aei Shirôzu & Shima, 1977; Kontyu 45 (4): 501; TL: Baguio City, Luzon Is, Philippines
  - Ypthima posticalis aeigma[sic]; [NHM card]
Ypthima parasakra Eliot, 1987 -. See [maps]
- Ypthima parasakra Eliot, 1987; Tyô to Ga 38 (1): 21, f. 1a-d, 2-5; TL: Nepal, Mustang District (28°40'N, 83°35'E), Larjung
- Y. p. parasakra Nepal, Bhutan
  - Ypthima parasakra parasakra; Huang, 2003, Neue Ent. Nachr. 55: 52 (note); Huang & Wu, 2003, Neue Ent. Nachr. 55: 124
  Misidentified/misapplied:
  
  Ypthima methorina; Lee, 1982, in Huang, Insects of Xizang, 2: 135, No. 23 (66) (missid.)
- Y. p. menpae Huang, 1999 SE.Tibet
  - Ypthima parasakra menpae; Huang, 2003, Neue Ent. Nachr. 55: 52 (note)
- Y. p. mabiloa Huang, 2003 Yunnan
  - Ypthima parasakra mabiloa Huang, 2003; Neue Ent. Nachr. 55: 52, pl. 7, f. 3; TL: Xiongdang, Dulong valley, NW.Yunnan
Ypthima striata Hampson, 1888 S.India (Nilgiri Hills). See [maps]
- Ypthima striata Hampson, 1888; J. asiat. Soc. Bengal 57 (4): 349; TL: Nilgiri Hills
- Ypthima striata; Elwes & Edwards, 1893, Trans. Ent. Soc. Lond. 1893 (1): 34, pl. 1, f. 23-24; [BOR], 466
Ypthima yamanakai Sonan, 1938 Taiwan
- Ypthima yamanakai Sonan, 1938; Zephyrus 7 (4): 250, pl. 20, f. 8; TL: Formosa
- Ypthima yamanakai; [BOR], 472 (note); [MRS], 391; Takahashi, 2000, Trans. lepid. Soc. Japan 51 (1): 9
Ypthima zyzzomacula Chou & Li, 1994 Yunnan. See [maps]
- Ypthima zyzzomacula Chou & Li, 1994; in Chou, Monographia Rhopalocerum Sinensium 1-2: 759, 395, f. 30-31; TL: Gengma, Yunnan
Ypthima wenlungi Takahashi, 2007 S.Taiwan
- Ypthima wenlungi Takahashi, 2007; Trans. lepid. Soc. Japan 58 (3): 277, f. 1-4; TL: Feng Gang Shan (1400-1600m), Kaohsiung, Taiwan

Unknown or unplaced taxa

selinuntioides (Mell, 1942) (Ypthima); Arch. Naturgesch. (N.F.) 11: 261

Ypthima selinuntioides; [NHM card]

Papilo lysandra Cramer, [1780]; Uitl. Kapellen 4 (25-26a): pl. 293

Yphthima[sic] lysandra; Hewitson, 1865, Trans. ent. Soc. Lond. (3) 2 (4): 287

Ypthima praenobilia
neobilia (Murayama, 1980) (Ypthima); New Entomologist 29 (2): 51

Ypthima praenobilia; [NHM card]

anana Fruhstorfer, 1911²; in Seitz, Gross-Schmett. Erde 9: 293; TL: E.Celebes

Ypthima anana; [NHM card]

27.2.2019 (126)

References:

[AfrL] Mark C. Williams
Checklist of Afrotropical Papilionoidea and Hesperoidea; Compiled by Mark C. Williams, 7th ed. (2008) (April 2007); Afrototropical Butterflies; e-mail: Mark C. Williams
[BHL] The
Biodiversity Heritage Library; and on; Internet Archive; Note: The linking to the content of Internet Archive is somewhat delicate -- notify me when links break!
[CiNii]
http://ci.nii.ac.jp/
[Don Herbison-Evans] and Stella A. Crossley
Lepidoptera Larvae of Australia; e-mail: Don Herbison-Evans; e-mail: Don Herbison-Evans
[Enrique Garcia-Barros]
Enrique García-Barros; Dept. Biol. (Zool.); Univ. Autonoma de Madrid
[FLMNH] Florida Museum of Natural History
Bulletin of the Allyn Museum
[HBN] Francis Hemming, 1937
HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[Kedar Tokekar]
India
[LAMAS-2005] Gerardo Lamas
Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[LEPIFORUM]
Lepiforum; Bestimmung von Schmetterlingen (Lepidoptera) und ihren Präimaginalstadien
[MBE] Moths and Butteflies of Europe
Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[NHM] Natural History Museum
The Global Lepidoptera Names Index; Butterflies and Moths of the World; Generic Names and their Type-species
[Oleg Kosterin] Institue of Cytology & Genetics
Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Olli Vesikko]
http://koti.welho.com/ovesikko/
[Peter Payzant]
Canada; http://www.pbase.com/ppayzant/profile
[RAFFLES]
Bulletin of the Raffles Museum (1928-1960); Bulletin of the National Museum (1970); The Raffles Bulletin of Zoology (1988-2013)
[SZMN] Lepidoptera
Siberian Zoological Museum
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[Yutaka Inayoshi]
A Check List of Butterflies in Indo-China; Chiefly from THAILAND, LAOS & VIETNAM
[ZOBODAT] Oberösterreichische Landesmuseum
OÖ. Landesmuseum; ZOBODAT - Literatur
[maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[²] This may require parentheses or not. I don't have the necessary information for this taxon.

Some related literature:

Aoki & Uémura, 1982
Description of new species and new subspecies of South-East Asian diurnal Lepidoptera belonging to the family Satyridae, with some taxonomic notes Memoirs of the Tsukada Coll. No. 4: 66pp., 8pls
Aoki & Uémura, 1984
Studies on the genus Ypthima in Thailand Tokurana 6/7: 71-103
Bartel, 1905
Neue Aethiopische Rhopalocera des Kgl. Museums für Naturkunde in Berlin Novit. Zool. 12 : 129-152
Bremer & Grey, 1853
Beiträge zur Schmetterlings-Fauna des nördlichen China Schmett. N. China : 5-23
Butler, 1866
A list of the diurnal Lepidoptera recently collected by Mr. Whitely in Hakodadi (North Japan) J. Linn. Soc. Zool. Lond. 9 (34) : 50-58
Butler, 1871
Descriptions of five new species, and a new genus , of diurnal Lepidoptera, from Shanghai Trans. ent. Soc. Lond. 1871 (3) : 401-403
Butler, 1874
Descriptions of four new Asiatic Butterflies Cistula ent. 1 (9) : 235-236
Butler, 1876
On a Collection of Lepidoptera recently received from Abyssinia Ann. Mag. nat. Hist. (4) 18 (108) : 480-490
Butler, 1879
The butterflies of Malacca Trans. linn. Soc. Lond. (2) 1 (8) : 533-568, pl. 68-69
Butler, 1880
On new and little-known Butterflies from India Proc. zool. Soc. Lond. 1880 : 147-152, pl. 15
Butler, 1881
On a Collection of Butterflies from Nikko, Central Japan Ann. Mag. Nat. Hist. (5) 7 (38) : 132-140
Butler, 1882
Descriptions of new Species of Lepidoptera from Tenasserim Ann. Mag. Nat. Hist. (5) 10 : 372-376
Butler, 1883
The Lepidoptera collected during the recent expedition of H.M.S. "Challenger." Ann. Mag. nat. Hist. (5) 11 (66) : 402-428
Butler, 1883
On a Third Collection of Lepidoptera made by Mr. H. E. Hobson in Formosa Ann. Mag. nat. Hist. (5) 12 (67) : 50-52
Butler, 1883
On some Lepidoptera from the Victoria Nyanza Ann. Mag. nat. Hist. (5) 12 (68) : 101-107
Butler, 1883
On a collection of Indian Lepidoptera received from Lieut.-Colonel Charles Swinhoe; with numerous notes by the collector Proc. zool. Soc. Lond. 1883 (2) : 144-175, pl. 24
Butler, 1886
On a Collection of Lepidoptera made by Commander Alfred Carpenter, R. N., in Upper Burma, in the Winter of 1885-86 Ann. Mag. nat. Hist. (5) 18 (105) : 182-191
Butler, 1888
On the Lepidoptera received from Dr. Emin Pascha [from Equatorial Africa] Proc. Zool. Soc. Lond. 1888 : 56-85
Cantlie & Norman, 1959
Notes on the butterfly genus Ypthima J. Bombay nat. Hist. Soc. 56 (1) : 66-71
Carcasson, 1961
New and Little-known African Lepidoptera Occ. Pap. Coryndon meml Mus. (7) : 3-23, 5pls
[MRS]; Chou, 1994
Monografia Rhopalocerorum Sinensium 1-2: 1-854
[BMP]; Corbet & Pendlebury, 1992
The butterflies of the Malay Peninsula. Fourth Edition revised by J. N. Eliot with plates by Bernard D'Abrera Butterflies Malay Peninsula (4th edn) : 1-595, pl. 1-69
Cowan, 1956
Two unusual varieties of common Malayan Rhopalocera Bull. Raffles Mus. 27 : 39-40
Cramer, [1780]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 4 (25-26a): 1-28, pl. 289-304 (1780), : 247-252 (index)
[BAUR]; D'Abrera, 1977
Butterflies of the Australian Region (2nd edn) Butts. Aust. Region: 1-416
de Nicéville, 1889
On new and little-known Butterflies from the Indian Region, with Revision of the Genus Plesioneura of Felder and of Authors J. Bombay nat. Hist. Soc. 4 (3) : 163-194, pl. A-B
de Nicéville, 1895
On new and little-known Butterflies from the Indo-Malayan Region J. Bombay nat. Hist. Soc. 10 (1): 13-40, (2): 169-194
de Nicéville, 1897
On new or little-known Butterflies from the Indo- and Austro-Malayan Regions J. Asiat. Soc. Bengal 66 Pt.II (3) : 543-577, pl. 1-4
Distant, 1882
Undescribed Rhopalocera from the Malay Peninsula Ann. Mag. nat. Hist. (5) 9 (53) : 396-397
Distant, 1882
On some apparently undescribed Rhopalocera Ann. Mag. nat. Hist. (5) 10 (59) : 405-407
Doherty, 1891
The Butterflies of Sumba and Sumbawa, with some Account of the Island of Sumba J. Asiat. Soc. Bengal 60 Pt.II (2) : 141-197, pl. 2
Donovan, 1805
An Epitome of the Natural History of the Insects of New Holland, New Zealand, New Guinea, Otaheite and other Islands in the Indian, Sothern and Pacific Oceans ... Epitome nat. Hist. Ins. New Holland : pl. [1-41]
Westwood, 1851
The genera of diurnal Lepidoptera, comprising their generic characters, a notice of their habitats and transformations, and a catalogue of the species of each genus; illustrated with 86 plates by W. C. Hewitson Gen. diurn. Lep. (1): 1-18, pl. A, 1-4 (1-2) (1846), (1): 19-132, pl. 4*, 18*, 5-25, 28 (3-14) (1847), (1): 133-200, pl. 1*, 26, 27, 29 (15-23) (1848), (1): pl. 31-44 (15-23) (1848), (1): 201-242, pl. 30, 45-52, 56-58, 60-62, 64 (24-31) (1849), (1): 243-250 [?], (2): 243-326, pl. 53-55, 63, 65, 66 (32-38) (1850), (2): 327-466, pl. 59, 67-77 (39-50) (1851), (2): 467-534, 143, 144, pl. 78-80 (Index) (51-54) (1852)
Dubatolov & Lvovsky, 1997
What is true Ypthima motschulskyi (Lepidoptera, Satyridae)? Trans. lepid. Soc. Japan 48 (4) : 191-198
K. L. & L. E. Dunn, 1991
Review of Australian Butterflies: distribution, life history and taxonomy. Pushlished by authors, Melbourne Review Austr. Butts. (1-4):
Eliot, 1959
New or little known Butterflies from Malaya Bull. Br. Mus. nat. Hist. (Ent.) 7 (8) : 371-391, 7fgs., pl. 10
Eliot, 1967
The Sakra Moore, 1857 Section of the Satyrid Genus Ypthima Hübner Entomologist 100 (1246) : 49-61, 1+22figs, pl. 2
Eliot, 1987
On some butteflies from Nepal, Bhutan and Thailand Tyô to Ga 38 (1): 21-29
van Eecke, 1913
Fauna Simalurensis. Lepidoptera Rhopalocera, fam. Satyridae, Morphidae and Nymphalidae Notes Leyden Mus. 35 : 243-247
Elwes & Edwards, 1893
A revision of the genus Ypthima, with especial reference to the characters afforded by the male genitalia Trans. Ent. Soc. Lond. 1893 (1) : 1-54, pl. 1-3
Eschscholtz, 1821
Beschreibung neuer ausländischer Schmetterling in Kotzebue, Entdeck. Reise Süd-See 3 (App. 5) : 201-219 (11pls)
Evans, 1912
A list of Indian butterflies J. Bombay nat. Hist. Soc. 21 (2): 553-584, (3): 969-1008
Evans, 1923; Evans, 1924
The Identification of Indian Butterflies J. Bombay nat. Hist. Soc. 29 (1): 230-260, pl. 1-9, (2): 519-537, pl. 10-12, (3): 780-797, pl. 13-16, (4): 890-907, pl. 17-21
Fabricius, 1775
Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, Adiectis Synonymis, Locis, Descriptionibus, Observationibus Syst. Ent. : 1-832
Felder, 1868
Diagnose neuer von E. Baron v. Ransonnet in Vorder-Indien gesammelter Lepidopteren Verh. zool.-bot. Ges. Wien 18 (1-2) : 281-286
C. & R. Felder, 1863
Lepidoptera nova a Dr. Carolo Semper in Insulis Philippinis collecta diagnosibus. (3) Wien. ent. Monats. 7 (4) : 105-127
Forster, [1948]
Beitrage zur Kenntnis der ostasiatischen Ypthima-Arten Mitt. Münch. Ent. Ges. 34 (2) : 472-492, pl. 30-33
Fruhstorfer, 1902
Beitrag zur Kenntniss der Lepidopteren der Viti-Inseln Stettin ent. Ztg 63 (1) : 350-359 (1 July)
Fruhstorfer, 1908
Lepidopterologisches Pêle-Mêle. VI. Neue Rhopaloceren von Formosa Ent. Zs. 22 (29) : 118-119 (17 October)
Fruhstorfer, 1908
Neue ostasiatische Satyriden Ent. Zs. 22 (31) : 127 (31 October)
Fruhstorfer, 1911
4. Familie: Satyridae in Seitz, Gross-Schmett. Erde 9 : 285-288 (14 February, 1911), : 289-296 (15 February, 1911), : 297-304 (22 February, 1911), : 305-312 (23 February, 1911), : 313-328 (before 25 April, 1911), : 329-336 (before 2 May, 1911), : 337-344 (before 9 May, 1911), : 345-352 (before 23 May, 1911), : 353-360 (before 30 May, 1911), : 361-368 (before 7 June, 1911), : 369-376 (before 13 June, 1911), : 377-384 (before 20 June, 1911), : 385-392 (before 27 June, 1911), : 393-400 (24 July), : 401 (15 August)
Godart, [1824]
Encyclopédie Méthodique. Histoire naturelle Entomologie, ou histoire naturelle des crustacés, des arachnides et des insectes Encyclopédie Méthodique 9 (1): 3-328 (1819), (2): 329-828 ([1824])
Grose-Smith, 1887
Descriptions of eight new Species of Asiatic Butterflies Ann. Mag. nat. Hist. (5) 20 : 265-268
Grünberg, 1910
in Schultze (Ed), Zoologische und anthropologische Ergebnisse einer Forschungsreise in Westlichen und Zentralen Süd-Afrika. Lepidoptera Denkschr. med.-naturw. Ges. Jena 16 (4) : 91-146
Guérin-Méneville, 1843
in Souvenirs d'un voyage dans l'Inde execute de 1834 a 1839, Animaux Articulés in Delessert, Souvenirs Voy. Inde (2) : 33-98, pl. 11-27
Hagen, 1902
Schmetterlinge von den Mentawej-Inseln Abh. Senckenb. Nat. Ges. 20 : 309-340, pl. 1-2
Hampson, 1888
The Butterflies of the Nilgiri District, South India J. asiat. Soc. Bengal 57 (4) : 346-368 (1889)
Hemming, 1967
Generic names of the butterflies and their type-species (Lepidoptera: Rhopalocera) Bull. Br. Mus. (Ent.) Suppl. 9 : 5-509
Hewitson, 1865
A Monograph of the Genus Yphthima, with descriptions of two new Genera of diurnal Lepidoptera Trans. ent. Soc. Lond. (3) 2 (4) : 281-294, pl. 17-18
Holland, 1887
Notes upon small collection of Rhopalocera made by Rev. B. C. Henry in the Island of Hainan, together with descriptions of some apparently new species Trans. Amer. Ent. Soc. 14 : 111-124
Huang, 2001
Report of H. Huang's 2000 Expedition to SE. Tibet for Rhopalocera Neue Ent. Nachr. 51 : 65-152
Huang, 2003
A list of butterflies collected from Nujiang (Lou Tse Kiang) and Dulongiang, China with descriptions of new species, and revisional notes Neue Ent. Nachr. 55 : 3-114
Huang & Wu, 2003
New and little known Chinese butteflies in the collection of the Institue of Zoology, Academia Sinica, Beijing - 1 Neue Ent. Nachr. 55 : 115-143
Huang & Xue, 2004
A Contribution to the Butterfly Fauna of Southern Yunnan Neue Ent. Nachr. 57: 135-154, pl. 11
Hübner, [1813]; Hübner, 1818
Zuträge zur Sammlung exotischer Schmettlinge, Vol. 1 [1808-] 1818 Zuträge Samml. exot. Schmett. 1 : [1-3], 4-6, [7], 8-32, [33-40] (post 22nd December 1818), : pl. [1-2], f. 1-12 (1808 - 1809), : pl. [3-25], f. 13-146 (1809 - 1813), : pl. [26-35], f. 147-200 (1814 - 1818)
Kalis, 1933
Bijdrage tot de kennis van de Lepidoptera Rhopalocera van Nederlandsch-Indië Tijdschr. Ent. 76 (1-2) : 47-86
Kashiwai, 1982
A New Species of the Genus Ypthima HUBNER from Mindanao, the Philippines (Lepidoptera, Satyridae) Tyô to Ga 33 (1,2): 44-50
Kielland, 1982
Revision of the genus Ypthima in the Ethiopian region excluding Madagascar (Lepidoptera, Satyridae) Tijdschr. Ent. 125 (5) : 99-154
Kirby, 1871
A synonymic catalogue of the diurnal Lepidoptera Syn. Cat. diurn. Lep.: vii + 690pp : 1-690
Kirby, 1880
Catalogue of the Lepidoptera (Rhopalocera, Sphingidae, Castniidae and Uraniidae) in the Museum of the Science and Art, Dublin, with remarks of new or interesting species Proc. R. Dublin Soc. (2) 2: 292-340
Kollar, [1844]
Aufzählung und Beschreibung der von Freiherr C. v. Hügel auf seiner Reise durch Kaschmir und das Himaleygebirge gesammelten Insekten in Hügel, Kaschmir und das Reich der Siek 4 : 393-564, pl. 1-28
Korb & Bolshakov, 2011
[A catalogue of butterflies (Lepidoptera: Papilioformes) of the former USSR. Second edition, reformatted and updated] (in Russian) Eversmannia Suppl. 2 : 1-123
Kudrna, 1974
An annotated List of Japanese Butterflies Atalanta 5 : 92-120
[BK]; Larsen, 1996
The Butterflies of Kenya and their natural history Butterflies of Kenya
Lee, 1982
Rhopalocera in Huang, Insects of Xizang, 2: 127-155
Leech, 1891
New Species of Lepidoptera from China Entomologist 24 (Suppl.) : 1-6
Leech, 1891
New Species of Rhopalocera from Western China Entomologist 24 (Suppl.) : 57-61, 66-68
Leech, [1892]
Butterflies from China, Japan, and Corea Butts China Japan Corea (1): 1-296, pl. 1-28 (1892-1894), (2): 297-681, pl. 29-43 (1893-1894)
[RFEL]; Leley, 2016
Annotated catalogue of the insects of Russian Far East. Volume II. Lepidoptera Cat. ins. Russian Far East. 2 : 1-812
[BOW]; Lewis, 1974
Butterflies of the World
Linnaeus, 1763
in Johansson (Thesis), Centuria Insectorum Amoenitates Acad. 6 : 384-415
Marshall, 1882
Notes on Asiatic Butterflies, with Descriptions of some new Species Proc. zool. Soc. Lond. 1882 (4) : 758-761
Marshall & de Nicéville, 1883
The Butterflies of India, Burmah and Ceylon, a descriptive Handbook of all the known Species of rhopalocerous Lepidoptera inhabiting that Region, with Notices of allied Species occurring in the neighbouring Countries along the Border, with numerous Illustrations. Vol. 1 Butts India Burmah Ceylon 1 (1): 1-94, pl. 1-9 (1882), (2): 95-327, pl. 10-17 (1883)
Martin, 1913
Neue Rhopaloceren aus Celebes Dt. ent. Z. Iris 27 (2): 109, (3): 121-129, pl. 6-8
Martin, 1929
Die Tagfalter der Insel Celebes. (8) Satyriden Mitt. Münch. Ent. Ges. 19 (5-9) : 117-164
Matsumura, 1906
Neue Rhopaloceren Japans Annot. zool. japon. 6 (1) : 7-15, pl. 1
Matsumura, 1909
Die Danaiden und Satyriden Japans Ent. Zs. 23 (19) : 91-92
Matsumura, 1919
Thousand Insects of Japan. Additamenta 3 [Shin Nihon senchu zukai] Thous. Ins. Japan. Addit. 3 : 475-742, pl. 26-53
Matsumura, 1928
New Butterflies especially from the Kuriles Insecta Matsumurana 2 (4) : 191-201
Matsumura, 1929
Some new butterflies from Japan, Korea and Formosa Insecta Matsumurana 3 (4) : 139-142
Mell, 1942
Die Amathusiidae und Satyriden Süd (un Südost-) Chinas Arch. Naturgesch. (N.F.) 11 : 221-290, pl. 1-3
Ménétriés, 1859
Lépidoptères de la Sibérie orientale et en particulier des rives de l'Amour in Schrenck, Reise Forschungen Amur-Lande 2 (1) : 1-75, pl. 1-5
Miller & Miller, 1978
"East Indian" butterflies: notes and descriptions. 1. Satyridae and Danaidae Bull. Allyn Mus. 49: 23pp
Moore, 1857; Moore, [1858]
A Catalogue of the Lepidopterous Insects in the Museum of the Hon. East-India Company in Horsfield & Moore, Cat. lep. Ins. Mus. East India Coy 1: : 1-278, pl. 1-12,1a,2a,3a,4a,5a,6a
Moore, [1875]
Descriptions of new Asiatic Lepidoptera Proc. zool. Soc. Lond. 1874 (4) : 565-579, pl. 66-67
Moore, 1878
List of Lepidopterous Insects collected by the late R. Swinhoe in the Island of Hainan Proc. zool. Soc. Lond. 1878 (3) : 695-708
Moore, 1878
A list of the Lepidopterous Insects collected by Mr. Ossian Limborg in Upper Tenasserim, with Descriptions of new Species Proc. zool. Soc. Lond. 1878 (4) : 821-859, pl. 51-53 (1879)
Moore, 1881
Descriptions of new Asiatic diurnal Lepidoptera Trans. ent. Soc. Lond. 1881 (3) : 305-313
Moore, [1880]
The Lepidoptera of Ceylon Lepid. Ceylon 1 (1): 1-40, pl. 1-18 (Dec 1880), (2): 41-80, pl. 19-36 (Jan 1881), (3): 81-136, pl. 37-54 (Jun 1881), (4): 137-190, pl. 55-71 (Dec 1881)
Moore, 1882
List of the Lepidoptera collected by the Rev. J.H. Hocking, chiefly in the Kangra Discrict, N.W Hiamalaya; with Descriptions of new Genera and Species Proc. zool. Soc. Lond. 1882 (1) : 234-263, pl. 11-12
Moore, 1884
Descriptions of some new Asiatic diurnal Lepidoptera; chiefly from specimens contained in the Indian Museum, Calcutta J. asiat. Soc. Bengal 53 Pt.II (1) : 16-52
Moore, [1893]
Lepidoptera Indica. Rhopalocera. Family Nymphalidae. Sub-families Satyrinae (continued), Elymniinae, Amathusiinae, Nymphalinae (Group Charaxina) Lepidoptera Indica 2 (13): 1-32, pl. 95-100 (1893), (14): 33-64, pl. 101-107 (1893), (15): 65-88, pl. 108-114 (1893), (16): 89-112, pl. 115-122 (1893), (17): 113-136, pl. 123-130 (1894), (18): 137-160, pl. 131-138 (1894), (19): 161-176, pl. 139-146 (1894), (20): 177-192, pl. 147-154 (1895), (21): 193-208, pl. 155-162 (1895), (22): 209-232, pl. 163-170 (1895), (23): 233-248, pl. 171-180 (1895), (24): 249-274, pl. 181-190 ([1896])
Morishita & Inomata, 1998
New or little known butterflies from Nepal Trans. lepid. Soc. Japan 49 (1): 1-8
Murayama, 1961
An unrecorded and some aberrant butterflies from Formosa [in Japanese] Tyô to Ga 11 (4): 54-61
Murayama, 1969
Some satyrid-butterflies of Japan and Korea, with descriptions of a new species and two new races Tohoku Konchu Kenkyu 4 (2): 17-21, pl. 2
Norman, 1958
A new butterflie from Assam J. Bombay nat. Hist. Soc. 55 (1) : 180-181
Oberthür, 1884
Lépidoptères du Thibet Étud. d'Ent. 9 : 7-22, pl. 1-2
Oberthür, 1891
Nouveaux Lépidoptères d'Asie Étud. d'Ent. 15 : 7-25, pl. 1-3
Okano, 1962
A new subspecies of Ypthima argus Butler from Formosa (Lepidoptera, Satyridae) Tyô to Ga 12 (3): 52-53
Overlaet, 1954
Satyrides Africains nouveaux Lambillionea 54 : 41-44
Overlaet, 1955
Exploration du Parc National d l'Upemba Miss. de Witte 27 : 1-106
Pendlebury, 1933
Notes and new Records of Butterflies from the Malay Peninsula J. Fed. Malay St. Mus. 17 (2) : 377-401
Poujade, 1885
[no title] Bull. Soc. Ent. Fr. (6) 5 : x-xi [10-11], : xxiv-xxv [24-25], : xli-xlii [41-42], : lxvi [61], : xciv-xcv [94-95], : cxliii [143], : cli [151], : cc [200], : ccvii-ccviii [207-208], : ccxv-ccxvi [215-216]
Prittwitz, 1867
Lepidopterologisches Stettin ent. Ztg 28 (7-9): 257-277 (1867)
Rothschild, 1892
Notes on a Collection of Lepidoptera made by William Doherty in Southern Celebes during August and September 1891. Part I, Rhopalocera Dt. ent. Z. Iris 5 (2) : 429-442
Scudder, 1875
Historical Sketch of the Generic Names proposed for Butterflies: A Contribution to Systematic Nomenclature Proc. Amer. Acad. Arts Sci. 10 (2) : 91-293
Shelford, 1904
A List of the Butterflies of Borneo with Descriptions of New Species J. Straits Asiat. Soc. 41 : 81-111
Shima, 1988
Phylogenetic relationships of the genus Ypthima Hübner (Lepidoptera, Satyridae) [in Japanese] Spec. Bull. lepid. Soc. Japan (6): 69-81
Shirôzu & Shima, 1977
New species and subspecies of the genus Ypthima Hübner from Southeast Asia Kontyu 45 (4): 501-509
[EBW]; Smart, 1976
The Illustrated Encyclopedia of the Butterfly World Illust. Encyp. Butt. World: -
Snellen, 1891
Aanteekeningen over eene verzameling Lepidoptera, in October 1889 van het eiland Flores ontvangen Tijdschr. Ent. 34 : 229-256
Snellen, 1892
Beschrijving van eenige nieuwe Javaansche Dagvlinders Tijdschr. Ent. 35 (3/4) : 133-147
van Son, 1955
Butterflies of southern Africa. II. Nymphalidae: Danainae and Satyrinae Transv. Mus. Mem. 8: 1-166, 37pls
Sonan, 1938
Notes on some butterflies from Formosa. (5) Zephyrus 7 (4) : 250-257
South, 1913
A list of butterflies collected by Captain F.M. Bailey in western China, south-eastern Tibet, and the Mishimi hills, 1911 J. Bomb. nat. Hist. Soc. 22 (2): 345-365, (3): 598-615
Strand, 1909
Zur Kenntniss afrikanische Ypthima-Arten auf Grund des im Berliner Museum vorhandenen Material Ent. Zs. 23 : 112-114
Strand, 1913
Zoologische Ergebnisse der Expedition des Herrn G. Tessmann nach Sud-Kamerun und Spanisch-Guinea. Lepidoptera Archiv Naturg. 79 A (7) : 138-151
Strand, 1914
Lepidoptera aus Kamerun gesammelt von Herrn Leutnant von Rothkirch un Panthen Arch. Naturgesch. 80 A (2) : 139-161
Sugitani, 1932
Some new aberrant forms of Japanese and Formosan butterflies [in Japanese] Zephyrus 4 (1): 5-9, pl. 2, (?): 88-91
Swinhoe, 1885
On the Lepidoptera of Bombay and the Deccan. Part I-IV Proc. Zool. Soc. Lond. 1885 : 124-148, pl. 9 (I. Rhopalocera), : 287-307, pl. 20-21 (II. Heterocera), : 447-476, pl. 27-28 (III. Heterocera), : 852-886, pl. 56-57 (IV. Heterocera)
Swinhoe, 1886
On the Lepidoptera of Mhow, in Central India Proc. zool. Soc. Lond. 1886 (4) : 421-465, pl. 40-41
Swinhoe, 1889
On new Indian Lepidoptera, chiefly Heterocera Proc. zool. Soc. Lond. 1889 (4) : 396-431, pl. 43-44
Takahashi, 2000
A revision of the Ypthima sakra group (Lepidoptera, Satyridae) in Taiwan, China, with description of a new species Trans. lepid. Soc. Japan 51 (1) : 1-18
Takahashi, 2007
A new species of Ypthima (Lepidoptera, Nymphalidae, Satyrinae) from southern Taiwan Trans. lepid. Soc. Japan 58 (3) : 276-280
Talbot, 1947
Fauna of British India including Ceylon and Burma. (Danaidae, Satyridae, Amathusiidae, and Acraeidae) Fauna Br. Ind., Butt. (2nd ed.) 2: xv + 406pp, 2pls
[BRU]; Tuzov, Bogdanov, Devyatkin, Kaabak, Korolev, Murzin, Samodurov, Tarasov, 1997
Guide to the Butterflies of Russia and adjacent territories: Hesperiidae, Papilionidae, Pieridae, Satyridae Butts. Russia adj. terr. 1
Uemura, 1984
A List of the Ypthima Species (Lepidoptera, Satyridae) Described by MATSUMURA, with Lectotype Designations Tyô to Ga 35 (3): 135-137
Uémura & Monastyrskii, 2000
Description of two new specis of the genus Ypthima Hübner (Lepidoptera: Satyridae) from North and Central Vietnam Trans. Lepid. Soc. Japan 51 (2) : 150-156
Ungemach, 1932
Contribution à l'étude des Lépidoptères d'Abyssinie (Pt. 1, Rhopalocères) Mém. Soc. Sci. nat. phys. Maroc 32 : 1-122, 2pls
Vane-Wright & de Jong, 2003
The butterflies of Sulawesi: annotated checklist for a critical island faunda Zool. Verh. Leiden 343: 3-268, pl. 1-16
Watson, 1897
Notes on a collection of butterflies from the north Chin Hills and upper Chindwin District, Burma J. Bombay nat. Hist. Soc. 10 (4) : 634-687, pl. A.
Westwood, 1888
List of diurnal Lepidoptera collected in northern Celebes by Dr Hickson Trans. Ent. Soc. Lond. 1888 (3) : 467-475, pl. 12
[BIR]; Wynter-Blyth, 1957
Butterflies of the Indian Region (1982 Reprint)

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