Life Insecta Lepidoptera Ditrysia Papilionoidea Nymphalidae

The higher classification here is not independent work, butis mostly based on Niklas Wahlberg's ongoing work, most recentalignment is based on November 2005 version. See [Wahlberg].

• Cithaerias Hübner, [1819] — 5 spp.
• Dulcedo d'Almeida, 1951 — 1 sp.
• Haetera Fabricius, 1807 — 2 spp.
• Pierella Herrich-Schäffer, 1865 — 11 spp.
• Pseudohaetera Brown, 1943 — 2 spp.

• Cyllogenes Butler, 1868 Evening Browns — 5 spp.
• Melanitis Fabricius, 1807 Evening Browns — 12 spp.
• Parantirrhoea Wood-Mason, 1881 — 1 sp.
• Gnophodes Doubleday, [1849] Evening Browns — 3 spp.
• Manataria Kirby, [1904] — 1 sp.
• Aeropetes Billberg, 1820 — 1 sp.
• Paralethe van Son, 1955 — 1 sp.

• Satyrites Scudder, 1872

  • Satyrites Scudder, 1872; (Fossil)

  • Satyrites (Lethina); [Wahlberg]
• Pseudoneorina Nel & Descimon, 1986

  • Pseudoneorina Nel & Descimon, 1986; (Fossil)

  • Pseudoneorina (Lethina); [Wahlberg]
• Neorinopsis Butler, 1873

  • Neorinopsis Butler, 1873; (Fossil)

  • Neorinopsis (Lethina); [Wahlberg]
• Aphysoneura Karsch, 1894 Painted Ringlets — 2 spp.
• Lethe Hübner, [1819] Treebrowns — 116 spp.
• Enodia Hübner, [1819] — 3 spp.
• Satyrodes Scudder, 1875 — 2 spp.
• Neope Moore, [1866] — 19 spp.
• Ninguta Moore, [1892] — 1 sp.
• Ptychandra C. & R. Felder, 1861 — 9 spp.
• Mandarinia Leech, [1892] — 1 sp.
• Chonala Moore, 1893 — 3 spp.
• Kirinia Moore, 1893 — 5 spp.
• Lasiommata Westwood in Humphreys & Westwood, 1841 — 16 spp. 3 spp.
• Lopinga Moore, 1893 — 5 spp. 1 sp.
• Orinoma Gray, 1846 — 2 spp.
• Pararge Hübner, [1819] — 3 spp. 1 sp.
• Rhaphicera Butler, 1867 — 3 spp.
• Tatinga Moore, 1893 — 1 sp.
• Nosea Koiwaya, 1993 — 1 sp.

• Callarge Leech, [1892] — 2 spp.
• Penthema Doubleday, [1848] — 5 spp.
• Zethera Felder, 1861 — 6 spp.
• Ethope Moore, [1866] — 4 spp.
• Neorina Westwood, [1850] Owls — 5 spp.

• Elymnias Hübner, 1818 Palmflies — 45 spp.

• Bicyclus Kirby, 1871 Bush Browns — 84 spp.
• Bletogona C. & R. Felder, [1867] — 2 spp.
• Mycalesis Hübner, 1818 Bushbrowns — 97 spp.
• Lohora Moore, 1880 — 17 spp.
• Nirvanopsis Vane-Wright in Vane-Wright & de Jong, 2003 — 1 sp.
• Hallelesis Condamin, 1961 — 2 spp.
• Heteropsis Westwood, 1850 — 61 spp.
• Orsotriaena Wallengren, 1858 — 2 spp.

• Coelites Westwood, [1850] — 3 spp.
• Erites Westwood, [1851] — 5 spp.

• Acrophtalmia C. & R. Felder, 1861 pale ringlets — 3 spp.
• Acropolis Hemming, 1934 — 1 sp.
• Ragadia Westwood, [1851] — 9 spp.

• Coenonympha Hübner, [1819] — 32 spp. 4 spp.
• Lyela Swinhoe, 1908 — 3 spp.
• Triphysa Zeller, 1850 — 4 spp.
• Sinonympha Lee, 1974 — 1 sp.

• Tarsocera Butler, [1899] — 7 spp.
• Dingana van Son, 1955 — 7 spp.
• Serradinga Henning & Henning, 1996 — 3 spp.
• Dira Hübner, [1819] — 4 spp.
• Torynesis Butler, [1899] — 5 spp.

• Erebia Dalman, 1816 Nokiperhoset Arguses — 108 spp. 6 spp.
• Diaphanos Adams & Bernard, 1981 — 3 spp.
• Ianussiusa Pyrcz & Viloria in Lamas, 2004 — 1 sp.
• Idioneurula Strand, 1932 — 1 sp.
• Manerebia Staudinger, 1897 — 37 spp.
• Neomaniola Hayward, 1949 — 1 sp.
• Stuardosatyrus Herrera & Etcheverry, 1965 — 1 sp.
• Sabatoga Staudinger, 1897 — 1 sp.
• Tamania Pyrcz, 1995 — 1 sp.

• Oressinoma Doubleday, [1849] — 2 spp.
• Palaeonympha Butler, 1871 — 1 sp.
• Archeuptychia Forster, 1964 — 1 sp.
• Caenoptychia Le Cerf, 1919 — 1 sp.
• Capronnieria Forster, 1964 — 1 sp.
• Cercyeuptychia Miller & Emmel, 1971 — 1 sp.
• Chloreuptychia Forster, 1964 — 10 spp.
• Cissia Doubleday, 1848 — 15 spp.
• Coeruleotaygetis Forster, 1964 — 1 sp.
• Cyllopsis R. Felder, 1869 — 29 spp.
• Erichthodes Forster, 1964 — 5 spp.
• Euptychia Hübner, 1818 — 15 spp.
• Cepheuptychia Forster, 1964 — 4 spp.
• Pharneuptychia Forster, 1964 — 6 spp.
• Caeruleuptychia Forster, 1964 — 19 spp.
• Euptychoides Forster, 1964 — 10 spp.
• Forsterinaria Gray, 1973 — 24 spp.
• Guaianaza Freitas & Peña, 2006 — 1 sp.
• Godartiana Forster, 1964 — 2 spp.
• Harjesia Forster, 1964 — 5 spp.
• Hermeuptychia Forster, 1964 — 8 spp.
• Magneuptychia Forster, 1964 — 29 spp.
• Megeuptychia Forster, 1964 — 2 spp.
• Megisto Hübner, [1819] — 2 spp.
• Moneuptychia Forster, 1964 — 5 spp.
• Neonympha Hübner, 1818 — 5 spp.
• Paramacera Butler, 1868 — 4 spp.
• Paryphthimoides Forster, 1964 — 13 spp.
• Pindis R. Felder, 1869 — 2 spp.
• Praefaunula Forster, 1964 — 3 spp.
• Pseudeuptychia Forster, 1964 — 2 spp.
• Rareuptychia Forster, 1964 — 1 sp.
• Splendeuptychia Forster, 1964 — 23 spp.
• Taydebis Freitas, 2003 — 1 sp.
• Taygetis Hübner, [1819] — 27 spp.
• Satyrotaygetis Forster, 1964 — 1 sp.
• Pseudodebis Forster, 1964 — 5 spp.
• Taygetina Forster, 1964 — 1 sp.
• Parataygetis Forster, 1964 — 2 spp.
• Pareuptychia Forster, 1964 — 7 spp.
• Posttaygetis Forster, 1964 — 1 sp.
• Taygetomorpha Miller in Lamas, 2004 — 2 spp.
• Yphthimoides Forster, 1964 — 23 spp.
• Zischkaia Forster, 1964 — 4 spp.

• Setodocis Billberg, 1820 — 1 sp.

• Argynnina Butler, 1867 — 2 spp.
• Argyronympha Mathew, 1886 — 5 spp.
• Argyrophenga Doubleday, 1845 — 3 spp.
• Dodonidia Butler, 1884 — 1 sp.
• Erebiola Fereday, 1879 — 1 sp.
• Erycinidia Rothschild & Jordan, 1905 — 6 spp.
• Geitoneura Butler, 1867 — 3 spp.
• Harsiesis Fruhstorfer, 1911 — 3 spp.
• Heteronympha Wallengren, 1858 — 7 spp.
• Hyalodia Jordan, 1924 — 1 sp.
• Hypocysta Westwood, [1851] — 12 spp.
• Lamprolenis Godman & Salvin, 1880 — 1 sp.
• Nesoxenica Waterhouse & Lyell, 1914 — 1 sp.
• Oreixenica Waterhouse & Lyell, 1914 — 6 spp.
• Paratisiphone Watkins, 1928 — 1 sp.
• Percnodaimon Butler, 1876 — 1 sp.
• Platypthima Rothschild & Jordan, 1905 — 8 spp.
• Altiapa Parsons, 1986 — 4 spp.
• Tisiphone Hübner, [1819] — 2 spp.
• Zipaetis Hewitson, 1863 Catseyes — 3 spp.

• Argyrophorus Blanchard, 1852 — 1 sp.
• Auca Hayward, 1953 — 4 spp.
• Chillanella Herrera, 1966 — 1 sp.
• Cosmosatyrus C. & R. Felder, [1867] — 1 sp.
• Elina Blanchard, 1852 — 2 spp.
• Etcheverrius Herrera, 1965 — 2 spp.
• Faunula C. & R. Felder, [1867] — 2 spp.
• Haywardella Herrera, 1966 — 1 sp.
• Homoeonympha C. & R. Felder, [1867] — 4 spp.
• Nelia Hayward, 1953 — 2 spp.
• Neomaenas Wallengren, 1858 — 8 spp.
• Neosatyrus Wallengren, 1858 — 1 sp.
• Palmaris Herrera, 1965 — 4 spp.
• Pampasatyrus Hayward, 1953 — 9 spp.
• Pamperis Heimlich, 1959 — 1 sp.
• Punargentus Heimlich, 1963 — 2 spp.
• Quilaphoetosus Herrera, 1966 — 2 spp.
• Spinantenna Hayward, 1953 — 1 sp.
• Tetraphlebia C. & R. Felder, [1867] — 1 sp.

• Aphantopus Wallengren, 1853 — 3 spp. 1 sp.
• Cercyonis Scudder, 1875 — 4 spp.
• Hyponephele Muschamp, 1915 Meadowbrowns — 65 spp. 1 sp.
• Maniola Schrank, 1801 Meadowbrowns — 7 spp. 1 sp.
• Proterebia Roos & Arnscheid, 1980 — 1 sp.
• Pyronia Hübner, [1819] — 5 spp.

• Melanargia Meigen, 1828 — 25 spp. 1 sp.

• Altopedaliodes Forster, 1964 — 9 spp.
• Apexacuta Pyrcz, 2004 — 4 spp.
• Arhuaco Adams & Bernard, 1977 — 2 spp.
• Calisto Hübner, 1823 — 46 spp.
• Cheimas Thieme, 1907 — 1 sp.
• Corades Hewitson, 1849 — 19 spp.
• Daedalma Hewitson, 1858 — 8 spp.
• Dangond Adams & Bernard, 1979 — 1 sp.
• Drucina Butler, 1872 — 2 spp.
• Druphila Pyrcz, 2004 — 1 sp.
• Eretris Thieme, 1905 — 17 spp.
• Eteona Doubleday, 1848 — 1 sp.
• Foetterleia Viloria, 2004 — 1 sp.
• Junea Hemming, 1964 — 2 spp.
• Lasiophila C. & R. Felder, 1859 — 13 spp.
• Lymanopoda Westwood, [1851] — 54 spp.
• Mygona Thieme, 1907 — 4 spp.
• Neopedaliodes Viloria, Miller & Miller in Lamas, 2004 — 10 spp.
• Oxeoschistus Butler, 1867 — 11 spp.
• Panyapedaliodes Forster, 1964 — 14 spp.
• Paramo Adams & Bernard, 1977 — 1 sp.
• Parapedaliodes Forster, 1964 — 1 sp.
• Pedaliodes Butler, 1867 — 158 spp.
• Praepedaliodes Forster, 1964 — 4 spp.
• Antopedaliodes Forster, 1964 — 1 sp.
• Pherepedaliodes Forster, 1964 — 3 spp.
• Physcopedaliodes Forster, 1964 — 1 sp.
• Corderopedaliodes Forster, 1964 — 3 spp.
• Praepronophila Forster, 1964 — 3 spp.
• Proboscis Thieme, 1907 — 2 spp.
• Pronophila Doubleday, [1849] — 17 spp.
• Protopedaliodes Viloria & Pyrcz, 1994 — 3 spp.
• Pseudomaniola Röber, [1889] — 10 spp.
• Punapedaliodes Forster, 1964 — 2 spp.
• Redonda Adams & Bernard, 1981 — 2 spp.
• Sierrasteroma Adams & Bernard, 1977 — 1 sp.
• Steremnia Thieme, 1905 — 7 spp.
• Steroma Westwood, [1850] — 3 spp.
• Steromapedaliodes Forster, 1964 — 4 spp.
• Thiemeia Weymer, 1912 — 1 sp.

• Arethusana de Lesse, 1951 — 1 sp.
• Argestina Riley, 1923 — 7 spp.
• Aulocera Butler, 1867 Banded Satyrs — 10 spp.
• Berberia de Lesse, 1951 — 2 spp.
• Boeberia Prout, 1901 — 1 sp.
• Brintesia Fruhstorfer, 1911 — 1 sp.
• Callerebia Butler, 1867 — 12 spp.
• Cassionympha van Son, 1955 — 3 spp.
• Chazara Moore, 1893 — 10 spp.
• Coenyra Hewitson, 1865 — 3 spp.
• Coenyropsis van Son, 1958 — 3 spp.
• Davidina Oberthür, 1879 — 2 spp.
• Hipparchia Fabricius, 1807 — 33 spp. 1 sp.
• Kanetisa Moore, 1893 — 1 sp.
• Karanasa Moore, 1893 — 37 spp.
• Loxerebia Watkins, 1925 — 19 spp.
• Mashuna van Son, 1955 — 2 spp.
• Melampias Hübner, [1819] — 1 sp.
• Minois Hübner, [1819] — 4 spp.
• Neita van Son, 1955 — 6 spp.
• Neocoenyra Butler, [1886] — 15 spp.
• Neominois Scudder, 1875 — 1 sp.
• Oeneis Hübner, [1819] — 35 spp. 3 spp.
• Paralasa Moore, 1893 — 32 spp.
• Paroeneis Moore, 1893 — 6 spp.
• Physcaeneura Wallengren, 1857 — 5 spp.
• Pseudochazara de Lesse, 1951 — 33 spp.
• Pseudonympha Wallengren, 1857 — 15 spp.
• Paternympha Henning & Henning, 1997 — 2 spp.
• Satyrus Latreille, 1810 — 13 spp.
• Strabena Mabille, 1877 — 37 spp.
• Stygionympha van Son, 1955 — 9 spp.
• Ypthima Hübner, 1818 Rings Ringlets — 126 spp.
• Austroypthima Holloway, 1974 — 1 sp.
• Ypthimomorpha van Son, 1955 — 1 sp.

• Amphidecta Butler, 1867 — 3 spp.
• Gyrocheilus Butler, 1867 — 1 sp.
• Neorinella Martins Neto & al., 1993 — 1 sp.

• Dyndirus Capronnier, 1874

  • Dyndirus Capronnier, 1874; Ann. Soc. ent. Belg. 17: 32 (nom. nud.)

  • Dyndirus; [NL4A], 224; [Wahlberg]

24.3.2019 (2309)

If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.