Speyeria

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Goto:

Speyeria Scudder, 1872

Speyeria Scudder, 1872; 4th Ann. Rep. Peabody Acad. Sci. (1871): 44; TS: Papilio idalia Drury
Semnopsyche Scudder, 1875; Bull. Buffalo Soc. nat. Sci. 2: 238, 258; TS: Papilio diana Cramer
- = ; [NHM card]; [NL4A], 261; Pelham, 2008, J. Res. Lepid. 40: 306; Dunford, 2009, Insecta Mundi 90: 7; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [Wahlberg]
Subgenus Mesoacidalia (Acidalia) Reuss, 1926; Dt. ent. Zs. 1926 (1): 69; TS: Papilio aglaja Linnaeus
- = ; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [Wahlberg]
- = Argynnis ; [RFEL]
Subgenus Proacidalia (Acidalia) Reuss, 1926; Dt. ent. Zs. 1926 (1): 69; TS: Argynnis clara Blanchard
- = Mesoacidalia ; [NHM card]; [Wahlberg]
Neoacidalia Reuss, 1926; Dt. ent. Zs. 1926 (1): 69; TS: Papilio cybele Fabricius
- = ; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 306; Dunford, 2009, Insecta Mundi 90: 7

Subgenus Semnopsyche (Acidalia); Reuss, 1926, Dt. ent. Zs. 1926 (1): 69
Mesoacidalia (Heliconiini); [NHM card]
Speyeria (Heliconiini); [NHM card]
Speyeria (Argynnina); Pelham, 2008, J. Res. Lepid. 40: 306
Speyeria (Argynnini); [NL4A], 261; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643; [Wahlberg]
Subgenus Speyeria (Argynnis); [RFEL]

700x734 (~72Kb) USA: nr Crystal Mountain (47°13'26N 120°40'11W), Kittitas Co., WA, 11.7.2000, Photo © Markku Savela

600x503 (~56Kb) USA: Cape Blanco, (42°49'57N 124°32'50W ±1km), Curry Co. OR, 2.8.2001, Photo © Markku Savela

The exact identification of these species is still unknown, but tentatively assumed to belong into this group.

Speyeria aglaja (Linnaeus, 1758)

EU, Morocco, Iran - Siberia, AC, China, Korea, Japan. See [maps] Orvokkihopeatäplä Dark Green Fritillary Stor pärlemorfjäril Dukatpärlefjäril Großer Perlmutterfalter Le Grand Nacré Lunares de Piata Dostojka aglaja

Papilio aglaja Linnaeus, 1758; Syst. Nat. (Edn 10) 1: 481; TL: Sweden
Papilio pasithoe Linnaeus, 1767; Syst. Nat. (Edn 12) 1 (2): 755 (repl. for Papilio aglaja Linnaeus)
Papilio emilia Acerbi, 1802; Travels North Cape 2: 253, pl. 2, f. 1-2
- = ; [NHM card]
Papilio charlotta Haworth, 1802; Prodr. Lep. Brit.: 3
- = ; [NHM card]
jurassina (Ragonot, 1871); Rev neuchateloise: [?]
Argynnis aglaja ab. eridioides Pflümer, 1879; Stettin Ent. Ztg 40: 157
- = ; [NHM card]
Argynnis aglaja ab. aberrans Lampa, 1885; Ent. Tidskr. 6 (1): 20; TL: Sweden, Jmt., Åreskutan
- = ; [NHM card]
Argynnis aglaja ab. wimani Holmgren, 1888; Ent. Tidskr. 9: 103, f. 1
- = ; [NHM card]
arvernensis Bramson, 1890²; Tagfalter Eur. Caucasus: 72
- = ; [NHM card]
suffusa (Tutt, 1896) (Argynnis); Brit. Butts: 290
- = Speyeria aglaja jurassina ; [NHM card]
aurea (Tutt, 1896) (Argynnis); Brit. Butts: 291
- = ; [NHM card]
flavescens (Tutt, 1896) (Argynnis); Brit. Butts: 291
- = (fuscans); [NHM card]
pallida (Tutt, 1896) (Argynnis); Brit. Butts: 291
- = ; [NHM card]
fusca (Tutt, 1896) (Argynnis); Brit. Butts: 292
- = ; [NHM card]
Argynnis aglaja methana Fruhstorfer, 1908; Ent. Zs. 22 (39): 161; TL: Sierra de Guadarrama, Castilia
Argynnis aglaia v. basalis Matsumura, 1908; Ent. Zs. 22 (39): 158; TL: Hokkaido
Argynnis aglaja plutus Oberthür, 1909; Étud. Lépid. Comp. 3: 209 (Leech, 1983 pl. 22, f. 6 ♀); TL: W.China, Wa-ssu-kow
- = ; [NHM card]
Argynnis aglaja ab. hortensia Riel, 1911; Ann. Soc. Linn. Lyon (n.s.) 58: 187 [ Donzel?]
Argynnis aglaia ab. gutta Wileman, 1911; Entomologist 44: 55; TL: Yezo, Jozanke
- = ; [NHM card]
Argynnis aglaia ashretha Evans, 1912; J. Bombay nat. Hist. Soc. 21 (2): 558, 582; TL: Chitral, Ashreth Valley
Argynnis aglaja ab. viridiatra Strand, 1912; Ent. Zs. 25 (49): 253
- = ; [NHM card]
Argynnis aglaja yopala Fruhstorfer, 1912; in Seitz, Gross-Schmett. Erde 9: 515; TL: Chitral
Argynnis aglaja taldena Fruhstorfer, 1912; in Seitz, Gross-Schmett. Erde 9: 516; TL: Setchuan
?Argynnis aglaja appenninicola Verity, 1914; Boll. Soc. ent. ital. 45: 213, f. 4-5
Argynnis aglaia ab. molybdina Newnham, 1917; Entomologist 50: 230
- = ; [NHM card]
Argynnis aglaja r. locuples Verity, 1919; Ent. Rec. J. Var. 31: 195
Argynnis aglaja r. emilocuples Verity, 1919; Ent. Rec. J. Var. 31: 195
- = ; [NHM card]
Argynnis aglaia r. locupletata Verity, 1922; Ent. Rec. J. Var. 34: 15 (repl. Argynnys aglaja r. locuples Verity, 1919)
scotica (Watkins, 1923) (Argynnis); Entomologist 56: 105
Argynnis aglaia v. purpura Evans, 1924; J. Bombay nat. Hist. Soc. 30 (1): 89
Argynnis aglaja mirabilis de Sagarra, 1925; Butll. Inst. catal. Hist. nat. (2) 5 (1): 45; TL: Orihuela del Tremedal (Aragó)
Argynnis aglaja montesignum de Sagarra, 1926; Butll. Inst. catal. Hist. nat. (2) 6 (6-7): 132
valesinoides (Reuss, 1926) (Acidalia); Dt. ent. Zs. 1926 (1): 69; TL: Kashmir
- = Speyeria aglaja fortuna ; [NHM card]
Argynnis aglaja ab. hindenburgi Schuster von Forstner, 1928; Lepid. Rdsch. 2 (16): 157
- = ; [NHM card]
ovalis (Cabeau, 1930) (Argynnis); Lambillionea 30: 179
- = ; [NHM card]
Argynnis aglaja f. clorinda de Sagarra, 1930; Butll. Inst. catal. Hist. nat. (2) 10 (7): 115
Argynnis aglaja r. fuscans Verity, 1935; Entomologist 68: 190; TL: Alpi Carniche, Sappada, 1300m
Argynnis aglaja r. rubidior Verity, 1935; Entomologist 68: 190; TL: Lago Maggiore
Argynnis aglaja r. subvitatha Verity, 1935; Entomologist 68: 202; TL: Tianchan, Aksultal
linnaei (Hemming, 1942) (Mesoacidalia); Proc. R. ent. Soc. Lond. (B) 11 (11): 159
Argynnis aglaja f. ocellata Eisner, 1942; Zool. Meded. Leiden 24 (4): 123; TL: Krummhübel, Riesengebirge
Argynnis aglaja f. duplicata Eisner, 1942; Zool. Meded. Leiden 24 (4): 123; TL: Amata, Lettland
Argynnis aglaja kansuensis Eisner, 1942; Zool. Meded. Leiden 24 (4): 123; TL: Sining, Kansu, 2500m
Argynnis aglaja ab. smrzi Slaby, 1949; Acta Soc. ent. Cal. 45 (1-2): 68
- = ; [NHM card]
Mesoacidalia charlotta f. pluriradiata Verity, 1950; Le Farfalle diurn. d'Italia 4: 301
?rubescens (Verity, 1950) (Mesoacidalia); Le Farfalle diurn. d'Italia 4: 302
- = Speyeria aglaja appennicola ; [NHM card]
Argynnis aglaia ab. robnora Kershaw, 1952; Ent. Rec. J. Var. 64: 69, pl. 3, f. 15
- = ; [NHM card]
Argynnis aglaia ab. mitchelli Kershaw, 1952; Ent. Rec. J. Var. 64: 69, pl. 3, f. 16
- = ; [NHM card]
Mesoacidalia charlotta f. cadmeis Lempke, 1956; Tijdschr. Ent. 99 (4): 175
Mesoacidalia charlotta f. transversa Lempke, 1956; Tijdschr. Ent. 99 (4): 175
Mesoacidalia charlotta f. infraochracea Lempke, 1956; Tijdschr. Ent. 99 (4): 175
sinenigra (Aagesen, 1977) (Mesoacidalia); Lepidoptera Kbh. 3 (2-3): 48
- = ; [NHM card]

Speyeria aglaja, Lepiforum [LEPIFORUM]
Argynnis aglaja, Schmetterlinge der Schweiz [Heiner Ziegler]
Argynnis aglaja, Moths and Butterflies of Europe [MBE]
Argynnis aglaja, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Argynnis aglaja, Svenska fjärilar [SVF]
Argynnis aglaja, Europaische Schmetterlinge [Mario Maier]
Argynnis aglaja, Captain's European Butterfly Guide [Simon Coombes]

Mesoacidalia vitatha ashretha; [NHM card]
Argynnis aglaja appennicola[sic, recte appenninicola]; [NHM card]
Speyeria aglaja basalis; [NHM card]
Speyeria aglaja clorinda; [NHM card]
Speyeria aglaja duplicata; [NHM card]
Speyeria aglaja cadmeis; [NHM card]
Speyeria aglaja fuscans; [NHM card]
Speyeria aglaja hortensia; [NHM card]
Speyeria aglaja infraochracea; [NHM card]
Speyeria aglaja jurassina; [NHM card]
Speyeria aglaja kansuensis; [NHM card]
Speyeria aglaja linnaei; [NHM card]
Speyeria aglaja locupletata; [NHM card]
Speyeria aglaja methana; [NHM card]
Speyeria aglaja mirabilis; [NHM card]
Speyeria aglaja montesignum; [NHM card]
Speyeria aglaja ocellata; [NHM card]
Speyeria aglaja pluriradiata; [NHM card]
Speyeria aglaja purpura; [NHM card]
Speyeria aglaja rubidior; [NHM card]
Speyeria aglaja scotica; [NHM card]
Speyeria aglaja semilocuples[sic, recte emilocuples?]; [NHM card]
Speyeria aglaja taldena; [NHM card]
Speyeria aglaja transversa; [NHM card]
Speyeria aglaja subvitatha; [NHM card]
Speyeria aglaja yopala; [NHM card]
Argynnis aglaja ab. wimani; Fruhstorfer, 1906, Ent. Zs. 20 (3): 17
Argynnis aglaja; Grum-Grshimailo, 1890, in Romanoff, Mém. Lép. 4: 439
Mesoacidalia aglaja; [BOW]: pl. 3, f. 13-14; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 31
Mesoacidalia aglaia[sic, recte aglaja]; [EBW]
Argynnis aglaja; [BRU2]: 36, pl. 23, f. 1-15; [Otakar Kudrna]; Yakovlev, 2012, Nota lepid. 35 (1): 92
Argynnis (Speyeria) aglaja; [RFEL]
Speyeria aglaja; [MRS], 473; [NHM card]; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643

500x363 (~61Kb) upperside West down, Devo, UK 14/06/95, Photo © Simon Coombes	500x389 (~50Kb) underside West down, Devo, UK 14/06/95, Photo © Simon Coombes	700x544 (~41Kb) upperside Finland, Kangasala, Ruutana, 682:34, 8/1997, Photo © Tero Piirainen
780x696 (~49Kb) FINLAND: Ab: Suomusjärvi, 669:31, 29.6.1997, Photo © Markku Savela	500x667 (~41Kb) ova Finland, Sipoo 670:39, 22.7.1980, Photo © Pertti Pakkanen	550x371 (~71Kb) larva Central Europe 1990-2000, Photo © Michael Borsch
550x371 (~80Kb) pupa Central Europe 1990-2000, Photo © Michael Borsch	597x400 (~59Kb) upperside Central Europe 1990-2000, Photo © Michael Borsch	777x565 (~100Kb) upperside female Russia, Moscow area, 29.7.2008, Photo © D. Smirnov
551x607 (~112Kb) upperside female Russia, Moscow area, 29.7.2008, Photo © D. Smirnov

Larva on Viola verecunda, Polygonum suffultum [MRS], Viola bicolor, Bistorta major [BRU2], 36

S. a. aglaja SEU, CEU, Altai, Sayan, W.Siberia, S.Siberia
- Argynnnis aglaja aglaja; [BRU2], 36
- Mesoacidalia aglaja aglaja; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 31
S. a. vitatha (Moore, [1875]) Pamirs
- Argynnis vitatha Moore, [1875]; Proc. zool. Soc. Lond. 1874 (4): 568; TL: Cashmere
- ?Argynnis aglaja cannelata Peschke, 1934; Int. ent. Zs. 28: 431; TL: Kashmir
- Mesoacidalia vitatha; [EBW], (name only); [NHM card]
- Mesoacidalia aglaja vitatha; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 31
- Speyeria aglaja vitatha; [MRS], 473
- Argynnis aglaja vitatha; [BRU2], 36
S. a. fortuna (Janson, 1877) Japan
- Argynnis fortuna Janson, 1877; Cistula ent. 2: 154; TL: Japan
- Argynnis aglaja fortuna f. myonia Fruhstorfer, 1907; Int. ent. Zs. 1 (34): 257; TL: Nagasaki
  - = Speyeria aglaja fortuna ; [NHM card]
- Argynnis aglaia fortuna ab. shanae Kuwayama, 1921; Dobuts. Z. 33: 333, 336, f. 3; TL: Etoroff I.
  - = Speyeria aglaja fortuna ; [NHM card]
- Argynnis aglaia fortuna ab. wilemana Kuwayama, 1921; Dobuts. Z. 33: 334, 336, f. 4; TL: Sapporo
  - = Speyeria aglaja fortuna ; [NHM card]
- Argynnis aglaia myonia ab. isshikii Matsumura, 1928; Insecta Matsumurana 2 (4): 193; TL: Hokkaido
  - = Speyeria aglaja fortuna ; [NHM card]
- Speyeria aglaja fortuna; [NHM card]
- Mesoacidalia charlotta fortuna; Kudrna, 1974, Atalanta 5: 107
- Mesoacidalia aglaja fortuna; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 31
S. a. ottomana (Röber, 1896) Armenia, Talys, Kopet-Dagh
- Argynnis aglaja var. ottomana Röber, 1896; Ent. Nachr. 22 (6): 82
- Speyeria aglaja ottomana; [NHM card]
- Argynnis aglaja ottomana; [BRU2], 36
S. a. kenteana (Stichel, 1901) Transbaikalia, N.Ussuri, Amur
- Argynnis aglaja kenteana Stichel, 1901; Berl. ent. Z. 46: 23 (Stgr. i. l.)
- Argynnis aglaja graeseri Kardakoff, 1928; Ent. Mitt. 17 (4): 270, pl. 7, f. 7-8; TL: Ussuri
- Speyeria aglaja graeseri; [NHM card]
- Mesoacidalia aglaja kenteana; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 31
- Argynnis aglaja kenteana; [BRU2], 36; [RFEL]
S. a. bessa (Fruhstorfer, 1907)
- Argynnis aglaja bessa Fruhstorfer, 1907; Int. ent. Zs. 1 (34): 257; TL: Tse-Kou
- Speyeria aglaja bessa; [NHM card]; Huang, 2001
S. a. auxo (Jachontov, 1909) Caucasus, Transcaucasus
- Argynnis aglaja var. auxo Jachontov, 1909; Revue Russe Ent. 8: 287; TL: Caucasus
- Speyeria aglaja auxo; [NHM card]
- Mesoacidalia aglaja auxo; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 31
S. a. lyauteyi (Oberthür, 1920) Morocco (Middle Atlas)
- Argynnis lyauteyi Oberthür, 1920; Étud. Lépid. Comp. 17: 48, pl. D; TL: Azrou (Morocco)
- Argynnis lyauteyi, Moths and Butterflies of Europe [MBE]
- Mesoacidalia liauteyi[sic, recte lyauteyi]; [EBW], (name only)
- Mesoacidalia aglaja lyauteyi; [BMAT]: 50, pl. 15, f. 10-13
- Speyeria aglaja lyauteyi; [NHM card]
435x600 (~32Kb) upperside Morocco, Ifrane, Moyen Atlas, Photo © Michel Tarrier 600x435 (~33Kb) underside Morocco, Ifrane, Moyen Atlas, Photo © Michel Tarrier
S. a. matsumurai (Nakahara, 1926) Kuriles
- Argynnis aglaja matsumurai Nakahara, 1926; Ins. Inscit. Mens. 14: 48
- Argynnis aglaia chishimensis Matsumura, 1928; Insecta Matsumurana 2 (4): 192
- Argynnis aglaja matsumurai; [BRU2], 36
- Speyeria aglaja matsumurai; [NHM card]
S. a. tonnai (Matsumura, 1928) Sakhalin
- Argynnis aglaia var. sachalinensis Matsumura, 1911; J. Coll. Agr. Tohoku Imp. Univ. Sapporo 4 (1): 43 (preocc.)
  - = Speyeria aglaja matsumurai ; [NHM card]
- Argynnis aglaia[sic] tonnai ab. otaniana Matsumura, 1928; Insecta Matsumurana 2 (4): 192; TL: Saghalien
- Speyeria aglaja otaniana; [NHM card]
- Argynnis aglaja tonnai; [BRU2], 36
S. a. clavimacula (Matsumura, 1929) S.Ussuri
- Argynnis aglaia[sic] clavimacula Matsumura, 1929; Insecta Matsumurana 3 (4): 154; TL: Korea
- Speyeria aglaja clavimacula; [MRS], 473; [NHM card]
- Argynnis aglaja clavimacula; [BRU2], 36
S. a. excelsior (Rothschild, 1933) Morocco (Rif Mts.)
- Argynnis aglaia excelsior Rothschild, 1933; Novit. Zool. 38: 317; TL: Ketama (Morocco)
- Mesoacidalia aglaja excelsior; [BMAT]: 50, pl. 15, f. 14-17
- Speyeria aglaja excelsior; [NHM card]
- Argynnis lyauteyi excelsior;
600x435 (~46Kb) upperside Morocco, Djebel Tisirène, Rif, Photo © Michel Tarrier 600x435 (~29Kb) underside Morocco, Djebel Tisirène, Rif, Photo © Michel Tarrier
S. a. gigasvitatha (Verity, 1935) Tian-Shan, Ghissar, Darvaz, Alai, S.Altai
- Argynnis aglaja gigasvitatha Verity, 1935; Entomologist 68: 202; TL: Issyk kul
- Argynnis aglaja alaicola Verity, 1935; Entomologist 68: 202
- Speyeria aglaja alaicola; [NHM card]
- Speyeria aglaja gigasvitatha; [NHM card]
- Argynnis aglaja gigasvitatha; [BRU2], 36
S. a. boreas (Hemming, 1942) EU, Siberia, Far East, Kamchatka
- Argynnis aglaja v. borealis Strand, 1901; Nyt Mag. Naturv. 39 (1): 49 (preocc. Argynnis thore boralis Staudinger, 1861)
  - = Speyeria aglaja boreas ; [NHM card]
- boreas (Hemming, 1942) (Mesocacidalia); Proc. R. ent. Soc. Lond. (B) 11 (11): 160 (repl. Argynnis aglaja borealis Strand, 1901)
- Mesocacidalia aglaja boreas; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 31
- Argynnis aglaja borealis; [BRU2], 36; [RFEL]

Speyeria alexandra (Ménétriés, 1832) N.Iran, W.Kopet-Dagh, Asia Minor. See [maps]
- Argynnis alexandra Ménétriés, 1832; Cat. Rais.: 246; TL: [Zuvand Plateau, Talysh Mts., Azerbaijan]
- Mesoacidalia alexandra; [BOW]: pl. 198, f. 5
- Speyeria aglaja alexandra; [NHM card]
- Argynnis alexandra; [BRU2]: 36, pl. 22, f. 7-9
- Fabriciana alexandra; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 31
- Speyeria alexandra; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643
Larva on Viola , V. canina, V. riviniana, V. tricolor [SPRK]
Speyeria clara (Blanchard, [1844]) -. See [maps]
- Argynnis clara Blanchard, [1844]; in Jacquemont, Voy. Inde 4 (Zool.): 20, pl. 2, f. 2-3
- Mesoacidalia clara; [NHM card]
- Argynnis clara; de Nicéville, 1889, J. Bombay nat. Hist. Soc. 4 (3): 165, pl. A, f. 6 ♀
- Speyeria clara; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643
- S. c. clara Kashmir, NW.India
  - Mesoacidalia clara clara; Lang & Han, 2009, Atalanta 40 (3/4): 494
- S. c. clarina (Staudinger, 1901) E.Qinghai, SW.Gansu, W.Sichuna, E.Tibet
  - Argynnis clara v. clarina Staudinger, 1901; Cat. Lep. palaearct. Faunengeb. 1: 38; TL: Amdo
  - Mesoacidalia clara neoclara Chou, Yuan, Yin, Zhang & Chen, 2002; Entomotaxonomia 24 (1): (52-68); TL: Gansu
    
    = Mesoacidalia clara clarina ; Lang & Han, 2009, Atalanta 40 (3/4): 494
  - Mesoacidalia clara clarina; [NHM card]
- S. c. manis (Fruhstorfer, 1903) Sikkim, S.Tibet
  - Argynnis clara manis Fruhstorfer, 1903; Dt. ent. Z. Iris 16 (1): 41; TL: Sikkim
  - Argynnis claudia Fawcett, 1904; Proc. zool. Soc. Lond. 2 (1): 136; TL: Khamba Jong, Thibet
    
    = Mesoacidalia clara manis ; Lang & Han, 2009, Atalanta 40 (3/4): 494
  - Argynnis claudia; [NHM card]
  - Mesoacidalia claudia; [BOW]: pl. 196, f. 10
  - Mesoacidalia clara manis; [NHM card]; Lang & Han, 2009, Atalanta 40 (3/4): 494
- S. c. menba (Lang, 2009) Tibet
  - Mesoacidalia clara menba Lang, 2009; Atalanta 40 (3/4): 494; TL: Tibet, Milin Co., Namula Pass, 3800m
- S. c. tongtianensis (Lang, 2009) Qinghai
  - Mesocacidalia clara tongtianensis Lang, 2009; Atalanta 40 (3/4): 495; TL: Qinghai, Qumalai Co., Chuhuier River, 4500m
- S. c. kanga (Lang, 2010) Sichuan
  - Mesocacidalia clara kanga Lang, 2010; Atalanta 41: 221; TL: Sichuan, Kangding, Muge Cuo Lake

Speyeria diana (Cramer, [1777])

SE.USA. See [maps] Diana

Papilio diana Cramer, [1777]; Uitl. Kapellen 2 (9-16): 4, 148, pl. 98, f. D, E

Speyeria diana, Moth Photographers Group [MPG]
Speyeria diana, Butterflies of America [BOA]
Speyeria diana, Butterflies and Moths of North America [BAMONA]
Speyeria diana, Georgia Lepidoptera (USA) [GALEPS]

Semnopsyche diana; Dyar, 1903, Bull. U.S. nat. Mus. 52: 12
Speyeria diana; [NACL], #4449; [EBW]; [BOW]: pl. 15, f. 15-16; [NHM card]; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 307; Dunford, 2009, Insecta Mundi 90: 11; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643

940x735 (~121Kb) USA: Alabama, 4.6.2005, Photo © Vitaly Charny	1078x876 (~155Kb) USA: Alabama, 4.6.2005, Photo © Vitaly Charny	1203x883 (~192Kb) upperside USA: Alabama, 4.6.2005, Photo © Vitaly Charny
1321x1137 (~214Kb) underside USA: Alabama, 4.6.2005, Photo © Vitaly Charny	1267x989 (~137Kb) USA: Alabama, 4.6.2005, Photo © Vitaly Charny

Larva on Viola papilionacea, V. cucullata, V. cornuta, V. sororia Dunford, 2009, Insecta Mundi 90: 12

Speyeria cybele (Fabricius, 1775)

USA. See [maps] Great Spangled Fritillary

Papilio cybele Fabricius, 1775; Syst. Ent.: 516

Speyeria cybele, Moth Photographers Group [MPG]
Speyeria cybele cybele, Butterflies of America [BOA]
Speyeria cybele, Butterflies and Moths of North America [BAMONA]

Argynnis cybele; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
Speyeria cybele; [NACL], #4450; [EBW]; [BOW]: pl. 15, f. 14, 17; [NHM card]; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 307; Dunford, 2009, Insecta Mundi 90: 13; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643

1024x761 (~81Kb) upperside USA: Taneum Creek, Wenatchee National Forest, (47°06'24N 120°54'44W), Kittitas Co., WA, 29.7.1999, Photo © Markku Savela	700x632 (~45Kb) underside USA: Taneum Creek, Wenatchee National Forest, (47°06'24N 120°54'44W), Kittitas Co., WA, 29.7.1999, Photo © Markku Savela	1000x790 (~91Kb) upperside male USA: nr Crystal Mountain / FR9726 (47°14'35N 120°40'41W), Kittitas Co., WA, 11.7.2000, Photo © Markku Savela
800x639 (~62Kb) underside male USA: nr Crystal Mountain / FR9726 (47°14'35N 120°40'41W), Kittitas Co., WA, 11.7.2000, Photo © Markku Savela	700x986 (~93Kb) female USA: Mima Mounds Natural Area Preserve, Thurston Co., WA, 13.8.2001, Photo © Markku Savela	1300x909 (~199Kb) upperside USA: Alabama, 26.5.2001, Photo © Vitaly Charny
1300x1065 (~179Kb) underside USA: Alabama, 26.5.2001, Photo © Vitaly Charny

Larva on Viola rotundifolia, V. papilionacea, V. palustris, V. adunca, Viola adunca var. bellidifolia, V. sororia, V. canadensis Dunford, 2009, Insecta Mundi 90: 14

S. c. cybele
- Papilio daphnis Cramer, [1775]; Uitl. Kapellen 1 (1-7): 89, pl. 57, f. E, F (preocc. Papilio daphnis [Schiffermüller], 1775)
  - = Speyeria cybele cybele ; Pelham, 2008, J. Res. Lepid. 40: 307
  - = Speyeria cybele ; [NHM card]; Dunford, 2009, Insecta Mundi 90: 13
- Argynnis cybele ab. ♂ baal Strecker, 1878; Synon. Cat. Macroplep.: 111
  - = Speyeria cybele cybele ; Pelham, 2008, J. Res. Lepid. 40: 307
  - = Speyeria cybele ; [NHM card]; Dunford, 2009, Insecta Mundi 90: 13
- Speyeria cybele cybele; Pelham, 2008, J. Res. Lepid. 40: 307
S. c. leto (Behr, 1862) California, Alberta, Wyoming, ...
- Argynnis leto Behr, 1862; Proc. Cal. Acad. Sci. 2: 173; TL: California
- Argynnis leto var. letis Wright, 1905; Butts. West Coast US: 52; TL: Northern California
  - = Speyeria cybele leto ; Pelham, 2008, J. Res. Lepid. 40: 308
- Acidalia (Semnopsyche) valesinoides-alba [= valesinoidesalba] Reuss, 1926; Dt. ent. Zs. 1926 (1): 69
  - = ?Speyeria cybele leto ; Pelham, 2008, J. Res. Lepid. 40: 308
  - = ?Speyeria nokomis nokomis ; Pelham, 2008, J. Res. Lepid. 40: 310
  - = Speyeria nokomis ; Dunford, 2009, Insecta Mundi 90: 15
- Argynnis leto tr.f. lethe Gunder, 1934; Can. Ent. 66 (6): 125; TL: Big Horn Mts, Wyoming
- Speyeria cybele leto, Butterflies of America [BOA]
- Speyeria leto lethe; [NHM card]
- Speyeria leto letis; [NHM card]
- Speyeria nokomis valesinoalba[sic, recte valesinoidesalba]; [NHM card]
- Speyeria leto valesinoidesalba; [NHM card]
- Argynnis leto; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
- Speyeria leto; [NHM card]
- Speyeria cybele leto; [NACL], #4450h; Pelham, 2008, J. Res. Lepid. 40: 307
S. c. carpenterii (Edwards, 1876) New Mexico
- Argynnis carpenterii Edwards, 1876; Trans. amer. ent. Soc. 5: 204; TL: New Mexico
- Speyeria cybele carpenterii, Butterflies of America [BOA]
- Argynnis cybele carpenterii; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
- Speyeria cybele carpenterii; [NACL], #4450d; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 307
S. c. charlottii (Barnes, 1897) Colorado
- Argynnis charlottii Barnes, 1897; Can. Ent. 29 (2): 39; TL: Glenwood Spring, Colorado
- Speyeria cybele charlottii, Butterflies of America [BOA]
- Speyeria cybele charlottii; [NACL], #4450e; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 307
S. c. krautwurmi (Holland, 1931) Michigan, ...
- Argynnis cybele var. krautwurmi Holland, 1931; Ann. Carnegie Mus. 20 (2): 255; TL: Les Cheneaux, Upper Peninsula of Michigan
- Speyeria cybele krautwurmi, Butterflies of America [BOA]
- Speyeria cybele krautwurmi; [NACL], #4450b; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 307
S. c. novascotiae (McDunnough, 1935) Nova Scotia
- Argynnis cybele novascotiae McDunnough, 1935; Can. ent. 67: 18; TL: White Point Beach, Queens Co., Nova Scotia
- Speyeria cybele novascotiae, Butterflies of America [BOA]
- Speyeria cybele novascotiae; [NACL], #4450a; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 307
S. c. pseudocarpenteri (Chermock & Chermock, 1940) Manitoba, ...
- Argynnis cybele pseudocarpenteri Chermock & Chermock, 1940; Can. Ent. 72 (4): 82; TL: Sand Ridge, Manitoba
- Speyeria cybele pseudocarpenteri, Butterflies of America [BOA]
- Speyeria cybele pseudocarpenteri; [NACL], #4450c; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 307
S. c. letona dos Passos & Grey, 1945 Utah
- Speyeria cybele letona dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 1; TL: City Creek Canyon, Salt Lake City, Utah
- Speyeria cybele letona, Butterflies of America [BOA]
- Speyeria leto letona; [NHM card]
- Speyeria cybele letona; [NACL], #4450f; Pelham, 2008, J. Res. Lepid. 40: 307
S. c. pugetensis Chermock & Frechin, 1947 Washington
- Speyeria cybele pugetensis Chermock & Frechin, 1947; Pan-Pacific Ent. 23 (3): 111; TL: Stimson Creek, nr Belfair, Mason Co., Washington
- Speyeria cybele pugetensis, Butterflies of America [BOA]
- Speyeria leto pugetensis; [NHM card]
- Speyeria cybele pugetensis; [NACL], #4450g; Pelham, 2008, J. Res. Lepid. 40: 308
S. c. eileenae Emmel, Emmel & Mattoon, 1998 California
- Speyeria cybele eileenae Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (12): 144; TL: California, Humboldt Co
- Speyeria cybele eileenae, Butterflies of America [BOA]
- Speyeria cybele eileenae; Pelham, 2008, J. Res. Lepid. 40: 308

Speyeria aphrodite (Fabricius, 1787)

Canada - Kansas. See [maps] Aphrodite Fritillary

Papilio aphrodite Fabricius, 1787; Mantissa Insectorum 2: 62
?daphnis (Martyn)
- = ; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13

Speyeria aphrodite, Moth Photographers Group [MPG]
Speyeria aphrodite, Butterflies of America [BOA]
Speyeria aphrodite, Butterflies and Moths of North America [BAMONA]

Argynnis aphrodite; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
Speyeria aphrodite; [NACL], #4451; [BOW]: pl. 15, f. 5-6; [NHM card]; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 308; Dunford, 2009, Insecta Mundi 90: 14; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643

1000x1262 (~165Kb) underside USA: Pike National Forest, Sugar Creek on CR-67 (about 39°18'N 105°10'W), Douglas Co., CO, 29.7.2012, Photo © Markku Savela

1000x1358 (~194Kb) upperside USA: Pike National Forest, Sugar Creek on CR-67 (about 39°18'N 105°10'W), Douglas Co., CO, 29.7.2012, Photo © Markku Savela

Larva on Viola lanceolata, V. fimbriatula, V. nuttallii, V. papilionacea, V. nephrophylla, Viola primulifolia var. acuta, V. sagittata, V. sororia, V. tricolor, V. adunca Dunford, 2009, Insecta Mundi 90: 15

S. a. aphrodite New York, Massachusetts, Ohio, ...
- Argynnis cybele f. bartschi Reiff, 1910; Psyche 17 (6): 252; TL: in West Roxbury, Mass.
  - = Speyeria aphrodite aphrodite ; Pelham, 2008, J. Res. Lepid. 40: 308
  - = Speyeria aphrodite ; [NHM card]; Dunford, 2009, Insecta Mundi 90: 14
- Argynnis aphrodite ab. bakeri Clark, 1932; Bull. U.S. nat. Mus. 157: 276, pl. 19, f. 1-2; TL: Waynesburg, Ohio
  - = Speyeria aphrodite aphrodite ; Pelham, 2008, J. Res. Lepid. 40: 308
  - = Speyeria aphrodite ; [NHM card]; Dunford, 2009, Insecta Mundi 90: 14
- Speyeria aphrodite ab. hughi Clark & Clark, 1951; Smiths. Misc. Coll. 116 (7): 60
  - = Speyeria aphrodite aphrodite ; Pelham, 2008, J. Res. Lepid. 40: 308
- Speyeria aphrodite aphrodite; Pelham, 2008, J. Res. Lepid. 40: 308
S. a. alcestis (Edwards, [1877]) Illinois, Iow, Colorado, Michigan
- Argynnis alcestis Edwards, [1877]; Trans. amer. ent. Soc. 5: 289
- Argynnis alcestis ab. suffusa Wolcott, 1916; Ent. News, 27 (3): 97; TL: bog S of Grand Rapids, Michigan
  - = Speyeria aphrodite alcestis ; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 308
- Speyeria aphrodite alcestis, Butterflies of America [BOA]
- Argynnis aphrodite alcestis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
- Speyeria aphrodite alcestis; [NACL], #4451b; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 308
S. a. columbia (H. Edwards, 1877) N.British Columbia, ...
- Argynnis columbia H. Edwards, 1877; Proc. Cal. Acad. Sc.: [1]; TL: Lahache, nr Alaskan border of British Columbia
- Speyeria aphrodite columbia, Butterflies of America [BOA]
- Argynnis columbia; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
- Speyeria aphrodite colombia[sic, recte columbia]; [NHM card]
- Speyeria aphrodite columbia; [NACL], #4451e; Pelham, 2008, J. Res. Lepid. 40: 309
S. a. byblis (Barnes & Benjamin, 1926) Arizona, ...
- Dryas aphrodite r. byblis Barnes & Benjamin, 1926; Bull. South. Calif. Acad. Sci. 25 (3): 91; TL: White Mts., Arizona
- Speyeria aphrodite byblis, Butterflies of America [BOA]
- Speyeria aphrodite byblis; [NACL], #4451g; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 309
S. a. winni (Gunder, 1932) Quebec
- Argynnis aphrodite r. winni Gunder, 1932; Can. Ent. 64 (12): 278; TL: St. Calixte de Kilkenny, Quebec
- Speyeria aphrodite winni, Butterflies of America [BOA]
- Speyeria aphrodite winni; [NACL], #4451a; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 309
S. a. whitehousei (Gunder, 1932) British Columbia
- Argynnis whitehousei Gunder, 1932; Can. Ent. 64 (12): 279; TL: Jafrray, B.C.
- Speyeria aphrodite whitehousei, Butterflies of America [BOA]
- Speyeria aphrodite whitehousei; [NACL], #4451d; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 309
S. a. ethne (Hemming, 1933)
- Argynnis ethne Hemming, 1933; Entomologist 66 (12): 276 (repl. Argynnis cypris Edwards, 1886)
- Argynnis cypris Edwards, 1886; Can. Ent. 18 (4): 62 (preocc. Argynnis cypris Meigen, 1828)
  - = Speyeria aphrodite ethne ; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 309
- Speyeria aphrodite ethne, Butterflies of America [BOA]
- Speyeria aphrodite ethne; [NACL], #4451f; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 309
S. a. manitoba (Chermock & Chermock, 1940) Manitoba, ...
- Argynnis manitoba Chermock & Chermock, 1940; Can. Ent. 72 (4): 83; TL: Sand Ridge, Manitoba
  - = Speyeria aphrodite mayae ; [NHM card]
- Argynnis aphrodite cypris tr.f. mayae Gunder, 1932; Can. Ent. 64 (12): 279; TL: Sand Ridge, Bener Dam Lake, nr Kelwood, Manitoba
  - = Speyeria aphrodite manitoba ; Pelham, 2008, J. Res. Lepid. 40: 309
- Speyeria (Semnopsyche) aphrodite mayae dos Passos & Grey, 1947; Amer. Mus. Novit. 1370: 8; TL: Sand Ridge, Bener Dam Lake, nr Kelwood, Manitoba
  - = Speyeria aphrodite manitoba ; Pelham, 2008, J. Res. Lepid. 40: 309
- Speyeria aphrodite manitoba, Butterflies of America [BOA]
- Speyeria aphrodite mayae; [NHM card]
- Speyeria aphrodite manitoba; [NACL], #4451c; Pelham, 2008, J. Res. Lepid. 40: 309
S. a. cullasaja Gatrelle, 1998 North Carolina, ...
- Speyeria aphrodite cullasaja Gatrelle, 1998; Taxonomic Rep. 1 (3): 2; TL: trail to Scaly Mtn, 4200', Macon Co., North Carolina
- Speyeria aphrodite cullasaja, Butterflies of America [BOA]
- Speyeria aphrodite cullasaja; Pelham, 2008, J. Res. Lepid. 40: 308

Speyeria idalia (Drury, [1773]) E.USA. See [maps] Regal Fritillary
- Papilio idalia Drury, [1773]; Illust. Nat. Hist. Exot. Insects 1: index, 25, pl. 13, f. 1-3
- Speyeria idalia, Moth Photographers Group [MPG]
- Speyeria idalia idalia, Butterflies of America [BOA]
- Speyeria idalia, Butterflies and Moths of North America [BAMONA]
- Speyeria idalia; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13; [NACL], #4452; [EBW]; [BOW]: pl. 16, f. 2; [NHM card]; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 309; Dunford, 2009, Insecta Mundi 90: 7; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643
Larva on Viola pedtifida, V. papilionacea, V. lanceolata, V. pedata, V. sagittata, V. sororia Dunford, 2009, Insecta Mundi 90: 10
- S. i. idalia New York, New Jersey, ...
  - Argynnis astarte Fisher, 1858; Proc. Acad. Nat. Sci. Philad. 10: 179 (preocc. Argynnis astarte Doubleday, 1847); TL: Nova Caesarea, Succasuma Plains, nr Schooley's Mtn, Morris Co.
    
    = Speyeria idalia idalia ; Pelham, 2008, J. Res. Lepid. 40: 309
    = Speyeria idalia ; [NHM card]; Dunford, 2009, Insecta Mundi 90: 7
  - Argynnis astaroth Fisher, 1859; Proc. Acad. Nat. Sci. Philad. 11: 352 (repl. Argynnis astarte Fisher, 1859); TL: Nova Caesarea, Succasuma Plains, nr Schooley's Mtn, Morris Co.
    
    = Speyeria idalia idalia ; Pelham, 2008, J. Res. Lepid. 40: 309
    = Speyeria idalia ; [NHM card]; Dunford, 2009, Insecta Mundi 90: 7
  - Argynnis idalia infumata Oberthür, 1912; Étud. Lépid. Comp. 6: 315, pl. 104, f. 969; TL: Newark, N.J.
    
    = Speyeria idalia idalia ; Pelham, 2008, J. Res. Lepid. 40: 310
    = Speyeria idalia ; Dunford, 2009, Insecta Mundi 90: 7
  - Argynnis idalia tr.f. dolli Gunder, 1927; Can. Ent. 59 (12): 286; TL: Richmond Hill, Long Island, N.Y.
    
    = Speyeria idalia idalia ; Pelham, 2008, J. Res. Lepid. 40: 310
    = Speyeria idalia ; Dunford, 2009, Insecta Mundi 90: 7
  - Argynnis idalia f. pallida Eisner, 1942; Zool. Meded. Leiden 24 (4): 123; TL: Decorah, Iowa
    
    = Speyeria idalia ; Dunford, 2009, Insecta Mundi 90: 7
  - Speyeria idalia dolli; [NHM card]
  - Speyeria idalia infumata; [NHM card]
  - Speyeria idalia pallida; [NHM card]
  - Speyeria idalia idalia; Pelham, 2008, J. Res. Lepid. 40: 309
- S. i. occidentalis Williams, 2002 Illinois, ...
  - Speyeria idalia occidentalis Williams, 2002; J. Lep. Soc. 55 (4): 146; TL: 1 mile SE of Crete, Illinois
  - Speyeria idalia occidentalis, Butterflies of America [BOA]
  - Speyeria idalia occidentalis; Pelham, 2008, J. Res. Lepid. 40: 310
Speyeria nokomis (Edwards, 1862) Utah, Arizona. See [maps] Nokomis Fritillary
- Argynnis nokomis Edwards, 1862; Proc. Acad. nat. Sci. Philad. 14: 221; TL: Mt. Sneffels, Ouray Co., Colorado
- Speyeria nokomis, Moth Photographers Group [MPG]
- Speyeria nokomis nokomis, Butterflies of America [BOA]
- Speyeria nokomis, Butterflies and Moths of North America [BAMONA]
- Argynnis nokomis; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 674; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
- Speyeria nokomis; [NACL], #4453; [EBW]; [BOW]: pl. 16, f. 8; [NHM card]; [Opler]; [NL4A], #2289; Pelham, 2008, J. Res. Lepid. 40: 310; Dunford, 2009, Insecta Mundi 90: 15; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643
Larva on Viola sororia Dunford, 2009, Insecta Mundi 90: 17
- S. n. nokomis California, New Mexico, ...
  - Argynnis nitocris var. nigrocaerulea Cockerell, 1900; Ent. News 11 (10): 622; TL: Beulah, Sapello Canon, New Mexico
    
    = Speyeria nokomis nokomis ; Pelham, 2008, J. Res. Lepid. 40: 310
  - Argynnis nitocris var. nigracaerulea[sic] ab. rufescens Cockerell, 1909; Ent. Rec. J. Var. 21 (7/8): 186; TL: Beulah, Sapello Cañon
    
    = Speyeria nokomis nigrocaerulea ; [NHM card]
    = Speyeria nokomis nokomis ; Pelham, 2008, J. Res. Lepid. 40: 310
  - Argynnis nitocris nigrocaerulea; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
  - Argynnis nokomis nigrocaerulea; [NHM card]
  - Speyeria nokomis nokomis; Pelham, 2008, J. Res. Lepid. 40: 310
- S. n. nitocris (Edwards, 1874) Arizona, ...
  - Argynnis nitocris Edwards, 1874; Trans. amer. ent. Soc. 5 (1): 15; TL: White Mtns, Arizona
  - Speyeria nokomis nitocris, Butterflies of America [BOA]
  - nitrocris[sic, recte nitocris]; dos Passos, 1964
  - Argynnis nitocris; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
  - Speyeria nitocris; [BOW]: pl. 16, f. 7; [NHM card]
  - Speyeria nokomis nitocris; [NACL], #4453a; Pelham, 2008, J. Res. Lepid. 40: 310
- S. n. coerulescens (Holland, 1900) Mexico
  - Argynnis nitocris var. coerulescens Holland, 1900; Ent. News 11 (1): 332; TL: Mexico, Chihuahua
  - Speyeria nokomis coerulescens, Butterflies of America [BOA]
  - Speyeria nokomis coerulescens, Butterflies of Southeastern Arizon [Bruce Walsh]
  - caerulescens[sic, recte coerulescens]; McDunnough, 1938
  - Argynnis nitocris var. coerulescens; Godman & Salvin, [1901], Biol. centr.-amer., Lep. Rhop. 2: 675, 3 pl. 112, f. 15-18
  - Argynnis nitocris caerulescens[sic, recte coerulescens]; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
  - Speyeria nokomis caerulescens[sic, recte coerulescens]; [NHM card]
  - Speyeria nokomis coerulescens; [NACL], #4453b; [NL4A], #2289a; Pelham, 2008, J. Res. Lepid. 40: 310
- S. n. apacheana (Skinner, 1918) California, ...
  - Argynnis apacheana Skinner, 1918; Ent. News 29 (2): 67; TL: near Independence, Inyo Co., California
  - Argynnis apacheana ab. hermosa Comstock, 1925; Bull. South. Calif. Acad. Sci. 24 (1): 3; TL: Round Valley, Inyo Co., California
    
    = Speyeria nokomis apacheana ; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 311
  - Speyeria nokomis apacheana, Butterflies of America [BOA]
  - Argynnis apacheana; Brown, 1983, J. Lep. Soc. 37 (1): 79-80
  - Speyeria nokomis apacheana; [NACL], #4453c; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 310
- S. n. wenona dos Passos & Grey, 1945 Mexico
  - Speyeria wenona dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: (1-17); TL: Cerro Potosi, Municipio de Galeana, Nuevo Leon, Mexico, 12000ft
  - Speyeria nokomis wenona, Butterflies of America [BOA]
  - Speyeria nokomis wenona; [NHM card]; [NL4A], #2289c; Pelham, 2008, J. Res. Lepid. 40: 448 (extralim.)
- S. n. melaena Mooser & Garcia, 1980 Mexico
  - nokomis melaena Mooser & Garcia, 1980; ; TL: Mexico, Aguascalienes
  - Speyeria nokomis melaena, Butterflies of America [BOA]
  - Speyeria nokomis melaena; [NL4A], #2289b
- S. n. carsonensis Austin, 1998 Nevada
  - Speyeria nokomis carsonensis Austin, 1998; Syst. W. N. Am. Butts. (46): 573
  - Speyeria nokomis carsonensis, Butterflies of America [BOA]
  - Speyeria nokomis carsonensis; Pelham, 2008, J. Res. Lepid. 40: 311
- S. n. tularosa Holland, 2010 New Mexico
  - Speyeria nokomis tularosa Holland, 2010; J. Res. Lepid. 42: (79-80)
  - Speyeria nokomis tularosa, Butterflies of America [BOA]
Speyeria edwardsii (Reakirt, 1866) Colorado, Utah, Nevada. See [maps] Edward's Fritillary
- Argynnis edwardsii Reakirt, 1866; Proc. Ent. Soc. Philad. 6 (1): 137; TL: Pikes Peak, Colorado
- Acidalia edwardsi montana Reuss, 1926; Dt. ent. Zs. 1926 (5): 439; TL: Montana
  - = ; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 311; Dunford, 2009, Insecta Mundi 90: 17
- Argynnis edwardsii tr.f. edonis Gunder, 1934; Can. Ent. 66 (6): 125; TL: Beaver City, Larimor Co., Colorado
  - = ; Pelham, 2008, J. Res. Lepid. 40: 311; Dunford, 2009, Insecta Mundi 90: 17
- Speyeria edwardsii, Moth Photographers Group [MPG]
- Speyeria edwardsii, Butterflies of America [BOA]
- Speyeria edwardsii, Butterflies and Moths of North America [BAMONA]
- Speyeria edwardsii edonis; [NHM card]
- Argynnis edwardsii; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
- Speyeria edwardsii; [NACL], #4454; [EBW]; [BOW]: pl. 15, f. 19; [NHM card]; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 311; Dunford, 2009, Insecta Mundi 90: 17; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643
Larva on Viola adunca, V. nuttallii Dunford, 2009, Insecta Mundi 90: 18

Speyeria coronis (Behr, 1864)

California. See [maps] Coronis Fritillary

Argynnis coronis Behr, 1864; Proc. ent. Soc. Philad. 3: 435; TL: California

Speyeria coronis, Moth Photographers Group [MPG]
Speyeria coronis coronis, Butterflies of America [BOA]
Speyeria coronis, Butterflies and Moths of North America [BAMONA]

Argynnis coronis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
Speyeria coronis; [NACL], #4455; [BOW]: pl. 15, f. 13; [NHM card]; [Opler]; [NL4A], #2288; Pelham, 2008, J. Res. Lepid. 40: 311; Dunford, 2009, Insecta Mundi 90: 18; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643

identification uncertain 600x834 (~64Kb) underside USA: FR 9705 x US Route 97, (47°18'38N 120°41'48W), Kittitas Co., WA, 14.8.2001, Photo © Markku Savela

identification uncertain 600x762 (~49Kb) USA: Table Mtn, Kittitas Co., WA, 30.7.1999, Photo © Markku Savela

identification uncertain 600x619 (~54Kb) underside USA: FR9708, Table Mtn, Kittitas Co., WA, 19.7.1999, Photo © Markku Savela

Larva on Viola beckwithii, Viola douglasii, V. nuttallii, V. purpurea, V. trinervata Dunford, 2009, Insecta Mundi 90: 19

S. c. coronis
- Argynnis californica Skinner, 1917; Ent. News 28 (7): 328; TL: California, Alma, Santa Clara Co.
  - = Speyeria coronis coronis ; Pelham, 2008, J. Res. Lepid. 40: 311
  - = Speyeria coronis ; [NHM card]; Dunford, 2009, Insecta Mundi 90: 18
- Speyeria coronis coronis; Pelham, 2008, J. Res. Lepid. 40: 311
S. c. hennei (Gunder, 1934) California, ...
- Argynnis hennei Gunder, 1934; Can. Ent. 66 (6): 126; TL: nr summit, 7700ft, Mt Pinos, Frazier Mtn Park, Santa Barbara Nat. Forest, Ventura Co., California
- Speyeria coronis hennei, Butterflies of America [BOA]
- Speyeria coronis hennei; [NACL], #4455a; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 311
S. c. semiramis (Edwards, 1886) WNA, California
- Argynnis semiramis Edwards, 1886; Can. Ent. 18 (4): 61; TL: California
- Argynnis semiramis tr.f. bernardensis Gunder, 1933; Can. Ent. 65 (8): 172; TL: California
  - = Speyeria coronis semiramis ; [NHM card]; [NL4A], #2288a; Pelham, 2008, J. Res. Lepid. 40: 311
- Speyeria coronis semiramis, Butterflies of America [BOA]
- Argynnis semiramis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
- Speyeria semiramis; [BOW]: pl. 16, f. 10
- Speyeria coronis semiramis; [NACL], #4455b; [NHM card]; [NL4A], #2288a; Pelham, 2008, J. Res. Lepid. 40: 311
S. c. simaetha dos Passos & Grey, 1945 Washington, ...
- Speyeria coronis simaetha dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 2; TL: Black Canyon, Cascade Mts., nr Brewster, Washington
- Speyeria coronis simaetha, Butterflies of America [BOA]
- Speyeria coronis simaetha; [NACL], #4455d; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 312
S. c. snyderi (Skinner, 1897) Utah
- Argynnis snyderi Skinner, 1897; Can. Ent. 29 (6): 154; TL: Salt Lake City, Utah
- Speyeria coronis snyderi, Butterflies of America [BOA]
- Argynnis snyderi; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
- Speyeria coronis snyderi; [NACL], #4455e; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 312
S. c. halcyone (Edwards, 1869) Colorado
- Argynnis halcyone Edwards, 1869; Butts. N. Amer. 1: 83; TL: Colorado
- Speyeria coronis halcyone, Butterflies of America [BOA]
- Argynnis halcyone; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
- Speyeria halcyone; [BOW]: pl. 15, f. 20
- Speyeria coronis halcyone; [NACL], #4455f; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 312

*S. coronis* or *S. zerene*?
1600x1171 (~287Kb) upperside USA: Utah, Uintah Co., Ashley National Park, start of FR-020 (40°40'35"N 109°29'14"W), 1.8.2012, Photo © Markku Savela	1600x1311 (~257Kb) underside USA: Utah, Uintah Co., Ashley National Park, start of FR-020 (40°40'35"N 109°29'14"W), 1.8.2012, Photo © Markku Savela	1300x1787 (~293Kb) USA: Utah, Uintah Co., Ashley National Park, start of FR-020 (40°40'35"N 109°29'14"W), 1.8.2012, Photo © Markku Savela
1500x1183 (~227Kb) USA: Utah, Uintah Co., Ashley National Park, start of FR-020 (40°40'35"N 109°29'14"W), 1.8.2012, Photo © Markku Savela	1200x1598 (~310Kb) upperside USA: Idaho, Fremont Co., on Hwy 20, Sheep Falls Road (44°11'39"N 111°26'39"W), 3.8.2012, Photo © Markku Savela	900x1001 (~128Kb) USA: Idaho, Fremont Co., on Hwy 20, Sheep Falls Road (44°11'39"N 111°26'39"W), 3.8.2012, Photo © Markku Savela
1200x1455 (~228Kb) USA: Idaho, Fremont Co., on Hwy 20, Sheep Falls Road (44°11'39"N 111°26'39"W), 3.8.2012, Photo © Markku Savela

Speyeria zerene (Boisduval, 1852)

California. See [maps] Zerene Fritillary

Argynnis zerene Boisduval, 1852; Ann. Soc. ent. Fr. (2) 10 (2): 303; TL: "Yosemite"? or Agua Fria, W of Mariposa

Speyeria zerene, Moth Photographers Group [MPG]
Speyeria zerene, Butterflies of America [BOA]
Speyeria zerene, Butterflies and Moths of North America [BAMONA]

Argynnis zerene; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
Speyeria zerene; [NACL], #4456; [EBW]; [BOW]: pl. 16, f. 9, 11; [NHM card]; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 313; Dunford, 2009, Insecta Mundi 90: 19; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643

identification uncertain 700x881 (~88Kb) underside USA: FR 9712 nr. Liberty, Lion Gulch, (47°15'47N 120°40'15W ±0.5km), Kittitas Co., WA, 14.8.2001, Photo © Markku Savela

identification uncertain 700x904 (~73Kb) USA: FR 9705 nr. Liberty and Lion Gulch, (47°17'49N 120°39'58W ±0.5km), Kittitas Co., WA, 14.8.2001, Photo © Markku Savela

identification uncertain 600x787 (~65Kb) underside USA: FR 9705 nr. US Route 97, (47°18'15N 120°41'24W ±0.5km), Kittitas Co., WA, 14.8.2001, Photo © Markku Savela

Larva on Viola adunca, Viola cuneata, V. lobata, V. nuttallii, V. psychodes, V. purpurea, V. beckwithii Dunford, 2009, Insecta Mundi 90: 21

S. z. zerene E.Oregon, California
- Argynnis zerene f. conchyliatus Comstock, 1925; Bull. South. Calif. Acad. Sci. 24 (3): 63; TL: Mt. Shasta, California
  - = Speyeria zerene zerene ; Pelham, 2008, J. Res. Lepid. 40: 313
- Argynnis zerene ab. shastaensis Comstock, 1925; Bull. South. Calif. Acad. Sci. 24 (3): 65; TL: Mt. Shasta, Siskiyo Co.
  - = Speyeria zerene conchyliatus ; [NHM card]
  - = Speyeria zerene zerene ; Pelham, 2008, J. Res. Lepid. 40: 313
- Speyeria zerene conchyliata; [NACL], #4456a
- Speyeria zerene conchyliatus; [NHM card]
- Speyeria zerene zerene; Pelham, 2008, J. Res. Lepid. 40: 313
S. z. monticola (Behr, 1863) California, ...
- Argynnis monticola Behr, 1863; Proc. Calif. Acad. Sci. 3 (1): 84; TL: Yosemite Valley
  - = Speyeria zerene ; Dunford, 2009, Insecta Mundi 90: 19; [NHM card]
- Speyeria zerene monticola, Butterflies of America [BOA]
- Argynnis monticola; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
- Speyeria zerene monticola; Pelham, 2008, J. Res. Lepid. 40: 313
S. z. behrensii (Edwards, 1869) California, ...
- Argynnis behrensii Edwards, 1869; Trans. amer. ent. Soc. 2 (4): 370; TL: Menocino, California
- Speyeria zerene behrensii, Butterflies of America [BOA]
- Argynnis behrensii; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
- Speyeria zerene behrensii; [NACL], #4456g; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 312
S. z. bremnerii (Edwards, 1872) inland (Vancouver Island - W.Washington - W.Oregon)
- Argynnis bremnerii Edwards, 1872; Trans. amer. ent. Soc. 4: 63; TL: San Juan I.
- Argynnis bremneri v. sordida Wright, 1905; Butts. West Coast US: 133; TL: Puget Sound, Washington
  - = Speyeria zerene bremnerii ; Pelham, 2008, J. Res. Lepid. 40: 312
- Speyeria zerene bremnerii, Butterflies of America [BOA]
- Speyeria zerene sordida; [NACL], #4456c; [NHM card]
- Argynnis bremnerii; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
- Speyeria zerene bremnerii; [NACL], #4456i; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 312
S. z. hippolyta (Edwards, 1879) coastal (California, W.Oregon, W.Washington)
- Argynnis hippolyta Edwards, 1879; Can. Ent. 11 (5): 81; TL: Oregon; Northern California
- Speyeria zerene hippolyta, Butterflies of America [BOA]
- Argynnis hippolyta; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
- Speyeria hippolyta; [BOW]: pl. 15, f. 21
- Speyeria zerene hippolyta; [NACL], #4456f; [NHM card]; McCorkle & Hammond, 1988, J. Lep. Soc. 42 (3): 184-195; Pelham, 2008, J. Res. Lepid. 40: 312
Larva on Viola adunca
S. z. platina (Skinner, 1897) Utah, Idaho
- Argynnis platina Skinner, 1897; Can. Ent. 29 (6): 154; TL: Ogden, Utan; Beaver Canon, Idaho
- Argynnis pfoutsi Gunder, 1933; Can. Ent. 65 (8): 171; TL: Mt. Loafer, nr Payson, Utah
  - = Speyeria zerene platina ; Grey, 1989, J. Lep. Soc. 43 (1): 3; Pelham, 2008, J. Res. Lepid. 40: 314
- Speyeria zerene platina, Butterflies of America [BOA]
- Argynnis platina; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
- Speyeria zerene pfoutsi; [NACL], #4456n; [NHM card]
- Speyeria zerene platina; [NACL], #4456m; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 314
S. z. malcolmi (Comstock, 1920) California
- Argynnis malcolmi Comstock, 1920; Southwest Sci. Bull. [1]: 4, pl. [1], f. 4-6; TL: Mammoth, California
- Argynnis montivaga r. malcolmi tr.f. sineargentatus Gunder, 1927; Can. Ent. 59 (12): 286; TL: Mammoth, Mono Co., California
  - = Speyeria zerene malcolmi ; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 313
- Speyeria zerene malcolmi, Butterflies of America [BOA]
- Speyeria zerene malcolmi; [NACL], #4456d; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 313
S. z. picta (McDunnough, 1924) British Columbia, ...
- Argynnis halcyone r. picta McDunnough, 1924; Can. ent. 56 (2): 43; TL: Aspen Grove, B. C.
- Argynnis garretti Gunder, 1932; Can. Ent. 64 (12): 282; TL: Cranbrook, B. C.
  - = Speyeria zerene picta ; [NACL], #4456j; Pelham, 2008, J. Res. Lepid. 40: 314
- Speyeria zerene picta, Butterflies of America [BOA]
- Speyeria zerene garretti; [NACL], #4456k; [NHM card]
- Speyeria zerene picta; [NACL], #4456j; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 314
S. z. gunderi (Comstock, 1925) California, ...
- Argynnis gunderi Comstock, 1925; Bull. South. Calif. Acad. Sci. 24 (3): 67; TL: Modoc Co., California
- Speyeria zerene cynna dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 4; TL: Humboldt Nat. Forest, Ruby Valley, Elko Co., NV
  - = Speyeria zerene gunderi ; Grey, 1989, J. Lep. Soc. 43 (1): 3; Pelham, 2008, J. Res. Lepid. 40: 313
- Speyeria zerene gunderi, Butterflies of America [BOA]
- Speyeria zerene cynna; [NACL], #4456o; [NHM card]
- Speyeria coronis gunderi; [NACL], #4455c; [NHM card]
- Speyeria zerene gunderi; Grey, 1989, J. Lep. Soc. 43 (1): 3; Pelham, 2008, J. Res. Lepid. 40: 313
S. z. myrtleae dos Passos & Grey, 1945 California
- Speyeria zerene myrtleae dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 3; TL: San Mateo, California
- Speyeria zerene myrtleae, Butterflies of America [BOA]
- Speyeria zerene myrtleae; [NACL], #4456h; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 313
S. z. sinope dos Passos & Grey, 1945 Colorado, ...
- Speyeria zerene sinope dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 4; TL: Estes Park area, Rocky Mt. National Park, Colorado
- Speyeria zerene sinope, Butterflies of America [BOA]
- Speyeria zerene sinope; [NACL], #4456l; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 314
S. z. gloriosa Moeck, 1957 Oregon, ...
- Speyeria zerene gloriosa Moeck, 1957; Milwaukee Ent. Soc., Special Paper: 21, 24; TL: Illinois River Valley, Josephine Co., SW.Oregon
- Speyeria zerene gloriosa, Butterflies of America [BOA]
- Speyeria zerene gloriosa; [NACL], #4456b; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 312
S. z. puntareyes Emmel, Emmel & Mattoon, 1998 California
- Speyeria zerene puntareyes Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (36): 451; TL: California, Marin Coo.; Point Reyes Peninsula
- Speyeria zerene puntareyes, Butterflies of America [BOA]
- Speyeria zerene puntareyes; Pelham, 2008, J. Res. Lepid. 40: 313
S. z. sonomensis Emmel, Emmel & Mattoon, 1998 California
- Speyeria zerene sonomensis Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (36): 455
- Speyeria zerene sonomensis, Butterflies of America [BOA]
- Speyeria zerene sonomensis; Pelham, 2008, J. Res. Lepid. 40: 313
S. z. sitka Hammond, Harry & McCorkle, 2001 Alaska
- Speyeria zerene sitka Hammond, Harry & McCorkle, 2001; Butts. B. C.: 274
- Speyeria zerene sitka, Butterflies of America [BOA]
- Speyeria zerene sitka; Pelham, 2008, J. Res. Lepid. 40: 312

Speyeria carolae (dos Passos & Grey, 1942) Nevada. See [maps]
- Argynnis coronis carolae dos Passos & Grey, 1942; Amer. Mus. Novit. 1214: 2; TL: Charleston Park, Clark Co., Nevada
- Speyeria carolae, Moth Photographers Group [MPG]
- Speyeria carolae, Butterflies of America [BOA]
- Speyeria carolae, Butterflies and Moths of North America [BAMONA]
- Speyeria zerene carolae; [NACL], #4456e; [NHM card]
- Speyeria carolae; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 312; Dunford, 2009, Insecta Mundi 90: 21; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643
Larva on Viola charlestonensis? Dunford, 2009, Insecta Mundi 90: 21
Speyeria callippe (Boisduval, 1852) -. See [maps] Callippe Fritillary
- Argynnis callippe Boisduval, 1852; Ann. Soc. ent. Fr. (2) 10 (2): 302; TL: San Francisco
- Speyeria callippe, Moth Photographers Group [MPG]
- Speyeria callippe, Butterflies of America [BOA]
- Speyeria callippe, Butterflies and Moths of North America [BAMONA]
- Argynnis callippe; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
- Speyeria callippe; [NACL], #4457; [BOW]: pl. 15, f. 11; [NHM card]; [Opler]; [NL4A], #2287; Pelham, 2008, J. Res. Lepid. 40: 314; Dunford, 2009, Insecta Mundi 90: 22; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643
Larva on Viola beckwithii, Viola douglasii, V. nuttallii, V. pedunculata, V. purpurea, V. purpurea quercetorum Dunford, 2009, Insecta Mundi 90: 23
- S. c. callippe California, ...
  - Speyeria callippe callippe; Pelham, 2008, J. Res. Lepid. 40: 314
- S. c. rupestris (Behr, 1863) California, ...
  - Argynnis rupestris Behr, 1863; Proc. Calif. Acad. Sci. 3 (1): 84; TL: California
  - ?irene (Edwards)
    
    = Argynnis rupestris ; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
  - Speyeria callippe rupestris, Butterflies of America [BOA]
  - Argynnis rupestris; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Speyeria callippe rupestris; [NACL], #4457e; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 315
- S. c. juba (Boisduval, 1869) California, Nevada
  - Argynnis juba Boisduval, 1869; Ann. Soc. ent. Belg. 12: 60
  - Argynnis laura Edwards, 1879; Can. Ent. 11 (3): 49; TL: Nevada
    
    = Speyeria callippe juba ; Pelham, 2008, J. Res. Lepid. 40: 315
  - Speyeria callippe sierra dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 5; TL: Gold Lake, Sierra Co., California
    
    = ?Speyeria callippe juba ; Grey, 1989, J. Lep. Soc. 43 (1): 4
    = Speyeria callippe juba ; Pelham, 2008, J. Res. Lepid. 40: 315
  - Speyeria callippe juba, Butterflies of America [BOA]
  - Speyeria callippe sierra; [NACL], #4457h; [NHM card]
  - Argynnis laura; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Speyeria callippe laura; [NACL], #4457g; [NHM card]
  - Speyeria callippe juba; [NACL], #4457f; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 315
- S. c. nevadensis (Edwards, 1870) Utah, Nevada, Wyoming
  - Argynnis nevadensis Edwards, 1870; Trans. amer. ent. Soc. 3 (1): 14; TL: valleys of the Sierra, nr Viriginia City
  - Speyeria callippe nevadensis, Butterflies of America [BOA]
  - Argynnis nevadensis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
  - Speyeria nevadensis; [BOW]: pl. 16, f. 6; [NHM card]
  - Speyeria callippe nevadensis; [NACL], #4457i; Pelham, 2008, J. Res. Lepid. 40: 316
- S. c. inornata (Edwards, 1872) California
  - Argynnis inornata Edwards, 1872; Trans. amer. ent. Soc. 4: 64; TL: "at Downieville, California"
    
    = Speyeria callippe juba ; [NHM card]
  - Speyeria callippe inornata, Butterflies of America [BOA]
  - Argynnis inornata; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Speyeria callippe inornata; Pelham, 2008, J. Res. Lepid. 40: 315
- S. c. meadii (Edwards, 1872) Colorado, ...
  - Argynnis meadii Edwards, 1872; Trans. amer. ent. Soc. 4: 69; TL: Turkey Creek Junction, Colorado
  - Argynnis nevadensis r. meadii tr.f. gerhardi Gunder, 1927; Can. Ent. 59 (12): 287; TL: Colorado
    
    = Speyeria nevadensis meadii ; [NHM card]
    = Speyeria callippe meadii ; Pelham, 2008, J. Res. Lepid. 40: 316
  - Speyeria callippe meadii, Butterflies of America [BOA]
  - Argynnis nevadensis meadii; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
  - Speyeria nevadensis meadii; [NHM card]
  - Speyeria callippe meadii; [NACL], #4457m; Pelham, 2008, J. Res. Lepid. 40: 316
- S. c. macaria (Edwards, 1877) California, Nevada
  - Argynnis macaria Edwards, 1877; Field & Forest 3 (5): 86; TL: havilah, California
  - Speyeria callippe macaria, Butterflies of America [BOA]
  - Speyeria macaria; [BOW]: pl. 16, f. 5
  - Speyeria callippe macaria; [NACL], #4457j; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 315
- S. c. liliana (H. Edwards, 1877) N.California
  - Argynnis liliana H. Edwards, 1877; Proc. Calif. Acad. Sci. 7 (12): 170; TL: nr St Helena, Napa Co.
  - Argynnis liliana var. baroni Edwards, 1881; Trans. Amer. Ent. Soc. 9: 3; TL: Northern California
    
    = Speyeria callippe liliana ; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 314
  - Speyeria callippe liliana, Butterflies of America [BOA]
  - Argynnis liliana; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
  - Argynnis liliana baroni; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
  - Speyeria callippe liliana; [NACL], #4457b; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 314
- S. c. laurina (Wright, 1905) California
  - Arynnis laurina Wright, 1905; Butts. West Coast US: 53; TL: Southern California
  - Argynnis wrightii Wright, 1905; Butts. West Coast US: 139
    
    = Speyeria callippe laurina (wrighti); [NHM card]
    = Speyeria callippe laurina ; Pelham, 2008, J. Res. Lepid. 40: 315 (nom. nud.)
  - Speyeria callippe laurina, Butterflies of America [BOA]
  - Speyeria callippe laurina; [NACL], #4457k; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 315
- S. c. semivirida (McDunnough, 1924) British Columbia
  - Argynnis callippe r. semivirida McDunnough, 1924; Can. ent. 56 (2): 42; TL: Aspen Grove, B.C.
  - Speyeria callippe semivirida, Butterflies of America [BOA]
  - Speyeria callippe semivirida; [NACL], #4457c; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 316
- S. c. calgariana (McDunnough, 1924) Alberta
  - callippe calgariana (McDunnough, 1924); Can. ent. 56 (2): 42; TL: Calgary .. Alta
  - Argynnis nevadensis r. calgariana McDunnough, 1924
  - Speyeria callippe calgariana, Butterflies of America [BOA]
  - Speyeria calgariana; [BOW]: pl. 15, f. 10
  - Speyeria nevadensis calgariana; [NHM card]
  - Speyeria callippe calgariana; [NACL], #4457o; Pelham, 2008, J. Res. Lepid. 40: 316
- S. c. comstocki (Gunder, 1925) California
  - Argynnis calippe[sic] var. comstocki Gunder, 1925; Ent. News 36: 8; TL: California
  - Argynnis comstocki tr.f. creelmani Gunder, 1934; Can. Ent. 66 (6): 127; TL: California
    
    = Speyeria callippe comstocki ; [NL4A], #2287a; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 315
  - Speyeria callippe comstocki, Butterflies of America [BOA]
  - Speyeria callippe comstocki; [NACL], #4457a; [NL4A], #2287a; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 315
- S. c. gallatini (McDunnough, 1929) Montana
  - Argynnis nevadenis var. gallatini McDunnough, 1929; Can. ent. 61 (5): 106; TL: Elkohn Ranch, Upper Gallatin Canyon, Montana
  - Speyeria callippe gallatini, Butterflies of America [BOA]
  - Speyeria nevadenis gallatini; [NHM card]
  - Speyeria callippe gallatini; [NACL], #4457n; Pelham, 2008, J. Res. Lepid. 40: 316
- S. c. elaine dos Passos & Grey, 1945 Oregon
  - Speyeria callippe elaine dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 5; TL: Butte Falls, Oregon
  - Speyeria callippe elaine, Butterflies of America [BOA]
  - Speyeria callippe elaine; [NACL], #4457d; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 316
- S. c. harmonia dos Passos & Grey, 1945 Nevada
  - Speyeria callippe harmonia dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 6; TL: Mt. Wheeler, Snake Range, nr. Utah border, NV, 8000ft
  - Speyeria callippe harmonia, Butterflies of America [BOA]
  - Speyeria nevadensis harmonia; [NHM card]
  - Speyeria callippe harmonia; [NACL], #4457l; Pelham, 2008, J. Res. Lepid. 40: 316
- S. c. shasta Emmel, Emmel & Mattoon, 1998 California
  - Speyeria callippe shasta Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (12): 149; TL: California, Shasta Co.
  - Speyeria callippe shasta, Butterflies of America [BOA]
  - Speyeria callippe shasta; Pelham, 2008, J. Res. Lepid. 40: 316
- S. c. adiasteoides Emmel, Emmel & Mattoon, 1998 California
  - Speyeria callippe adiasteoides Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (12): 150; TL: California, Mendocino Co.. Cahto Peak summit, 4000-4200ft
  - Speyeria callippe adiasteoides, Butterflies of America [BOA]
  - Speyeria callippe adiasteoides; Pelham, 2008, J. Res. Lepid. 40: 314
- S. c. hagemani Emmel, Emmel & Mattoon, 1998 California
  - Speyeria callippe hagemani Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (12): 150; TL: California, Shata Co., Shingletown, 3500'
  - Speyeria callippe hagemani, Butterflies of America [BOA]
  - Speyeria callippe hagemani; Pelham, 2008, J. Res. Lepid. 40: 316
- S. c. chilcotinensis Guppy & Shepard, 2001 British Columbia
  - Speyeria callippe chilcotinensis Guppy & Shepard, 2001; Butts. B.C.: 276; TL: BC, Riske Creek
  - Speyeria callippe chilcotinensis, Butterflies of America [BOA]
  - Speyeria callippe chilcotensis[sic, recte chilcotinensis]; Pelham, 2008, J. Res. Lepid. 40: 316
Speyeria egleis (Behr, 1862) -. See [maps] Egleis Fritillary
- Argynnis egleis Behr, 1862; Proc. Cal. Acad. Sci. 2: 174 (Boisd.)
- Argynnis egleis Boisduval, 1869; Ann. Soc. ent. Belg. 12: 59
- ?nenoquis (Reakirt)
  - = ; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
- Speyeria egleis, Moth Photographers Group [MPG]
- Speyeria egleis, Butterflies of America [BOA]
- Speyeria egleis, Butterflies and Moths of North America [BAMONA]
- Argynnis egleis Boisduval, 1869; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
- Speyeria egleis; [NACL], #4458; [NHM card]; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 317; Dunford, 2009, Insecta Mundi 90: 23; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643
Larva on Viola adunca, V. nuttallii, V. ocellata, Viola pupurea, V. purpurea integrifolia, V. purpurea venosa, V. walteri, Festuca ovina, Potentilla Dunford, 2009, Insecta Mundi 90: 24
- S. e. egleis California
  - Argynnis montivaga Behr, 1863; Proc. Calif. Acad. Sci. 3 (1): 84; TL: California
    
    = Speyeria egleis egleis ; Pelham, 2008, J. Res. Lepid. 40: 317
    = Speyeria egleis ; [NHM card]; Dunford, 2009, Insecta Mundi 90: 23
  - Argynnis egleis Boisduval, 1869; Ann. Soc. ent. Belg. 12: 59
    
    = Speyeria egleis egleis ; Pelham, 2008, J. Res. Lepid. 40: 317
  - Argynnis astarte Edwards, 1864; Proc. ent. Soc. Philad. 3: 434 (preocc. Arynnis astarte Doubleday, 1847)
    
    = Speyeria egleis egleis ; Pelham, 2008, J. Res. Lepid. 40: 317
    = Speyeria egleis ; [NHM card]; Dunford, 2009, Insecta Mundi 90: 23
  - Argynnis montivaga ab. mammothi Gunder, 1924; Ent. News 35 (5): 157, pl. 2, f. H, D
    
    = Speyeria egleis egleis ; Pelham, 2008, J. Res. Lepid. 40: 317
    = Speyeria egleis ; [NHM card]; Dunford, 2009, Insecta Mundi 90: 23
  - Argynnis montivaga tr.f. boharti Gunder, 1929; Bull. Brooklyn ent. Soc. 24 (5): 326, pl. 31, f. 5
    
    = Speyeria egleis egleis ; Pelham, 2008, J. Res. Lepid. 40: 317
    = Speyeria egleis ; [NHM card]; Dunford, 2009, Insecta Mundi 90: 23
  - Argynnis montivaga; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Speyeria egleis egleis; Pelham, 2008, J. Res. Lepid. 40: 317
- S. e. oweni (Edwards, 1892) S.Oregon - California
  - Argynnis oweni Edwards, 1892; Can. Ent. 24 (5): 105; TL: Mt. Shasta, Sikiyou Co., California
  - Speyeria egleis oweni, Butterflies of America [BOA]
  - Argynnys hippolyta oweni; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
  - Speyeria egleis oweni; [NACL], #4458e; [NHM card]; Hammond & Dornfeld, 1983, J. Lep. Soc. 37 (2): (note), f. 3; Pelham, 2008, J. Res. Lepid. 40: 317
- S. e. utahensis (Skinner, 1919) Utah
  - Argynnis utahensis Skinner, 1919; Ent. News 30 (8): 216; TL: City Creek Canyon, Salt Lake City, Utah
  - Speyeria egleis utahensis, Butterflies of America [BOA]
  - Speyeria egleis utahensis; [NACL], #4458i; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 318
- S. e. tehachapina (Comstock, 1920) California
  - Argynnis montivaga var. tehachapina Comstock, 1920; Southwest Sci. Bull. [1]: 6; TL: highest peak in Tehachapi Mts, California
  - Speyeria egleis tehachapina, Butterflies of America [BOA]
  - Speyeria egleis tehachapina; [NACL], #4458d; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 317
- S. e. macdunnoughi (Gunder, 1932) Montana
  - Argynnis macdunnoughi Gunder, 1932; Can. Ent. 64 (12): 280; TL: Elkhorn Ranch Resort, Gallatin Co., Montana
  - Speyeria egleis macdunnoughi, Butterflies of America [BOA]
  - Speyeria egleis macdunnoughi; [NACL], #4458g; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 318
- S. e. albrighti (Gunder, 1932) Montana
  - Argynnis albrighti Gunder, 1932; Can. Ent. 64 (12): 281; TL: Highwood Mts., Choteau Co., Montana
  - Speyeria egleis albrighti, Butterflies of America [BOA]
  - Speyeria egleis albrighti; [NACL], #4458h; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 318
- S. e. linda (dos Passos & Grey, 1942) Idaho
  - Argynnis utahensis linda dos Passos & Grey, 1942; Amer. Mus. Novit. 1214: 1; TL: Heyburn Peak, Sawtooth-Boise, ID, 9500-10000ft
  - Speyeria egleis linda, Butterflies of America [BOA]
  - Speyeria egleis linda; [NACL], #4458f; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 318
- S. e. secreta dos Passos & Grey, 1945 Colorado
  - Speyeria montiviga[sic] secreta dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 7; TL: Estes Park area, Rocky Mt. National Park, Colorado, 8000ft
  - Speyeria egleis secreta, Butterflies of America [BOA]
  - Speyeria egleis secreta; [NACL], #4458k; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 318
- S. e. toiyabe Howe, 1975 Nevada
  - Speyeria egleis toiyabe Howe, 1975; Butterflies of North America: 237; TL: Kingston Canyon, Toiyabe Range, Lander Co., Nevada
  - Speyeria egleis toiyabe, Butterflies of America [BOA]
  - Speyeria egleis toiyabe; [NACL], #4458j; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 318
- S. e. moecki Hammond & Dornfeld, 1983 Oregon
  - Speyeria egleis moecki Hammond & Dornfeld, 1983; J. Lep. Soc. 37 (2): 115, f. 2; TL: Skookum Mdw., Walker Rim, Klamath Co., Oregon
  - Speyeria egleis moecki, Butterflies of America [BOA]
  - Speyeria egleis moecki; Pelham, 2008, J. Res. Lepid. 40: 318
  Larva on Viola purpurea Hammond & Dornfeld, 1983, J. Lep. Soc. 37 (2): 120
- S. e. mattooni Emmel & Emmel, 1998 California
  - Speyeria egleis mattooni Emmel & Emmel, 1998; Syst. W. N. Am. Butts. (33): 438; TL: California, Siskiyou Co., Dry Lake Mtn, US FSR 40S01
  - Speyeria egleis mattooni, Butterflies of America [BOA]
  - Speyeria egleis mattooni; Pelham, 2008, J. Res. Lepid. 40: 317
- S. e. yolaboli Emmel & Emmel, 1998 California
  - Speyeria egleis yolaboli Emmel & Emmel, 1998; Syst. W. N. Am. Butts. (33): 439; TL: California, Tehama Co., South Yolla Bolly Mtn
  - Speyeria egleis yolaboli, Butterflies of America [BOA]
  - Speyeria egleis yolaboli; Pelham, 2008, J. Res. Lepid. 40: 317
- S. e. reidi Austin, 1998 Nevada
  - Speyeria egleis reidi Austin, 1998; Syst. W. N. Am. Butts. (46): 574; TL: Nevada, White Pine Co., Snake Range, Wheeler Peak Campground, 3033m
  - Speyeria egleis reidi, Butterflies of America [BOA]
  - Speyeria egleis reidi; Pelham, 2008, J. Res. Lepid. 40: 318
Speyeria adiaste (Edwards, 1864) California. See [maps] Unsilverered Fritillary Adiaste Fritillary
- Argynnis adiaste Edwards, 1864; Proc. ent. Soc. Philad. 3: 436; TL: California
- Speyeria adiaste, Moth Photographers Group [MPG]
- Speyeria adiaste adiaste, Butterflies of America [BOA]
- Speyeria adiaste, Butterflies and Moths of North America [BAMONA]
- Argynnis adraste; Kirby, 1871, 160 (missp.)
- ?Argynnis adiaste Behr, 1862; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
- Speyeria adiaste; [Opler]; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 318; Dunford, 2009, Insecta Mundi 90: 24; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643
Larva on Viola purpurea quercetorum, V. ocellata? Dunford, 2009, Insecta Mundi 90: 25
- S. a. adiaste
  - Argynnis adiante Boisduval, 1869; Ann. Soc. ent. Belg. 12: 61; TL: California
    
    = Speyeria adiaste ; [NHM card]
    = Speyeria adiaste adiaste ; Pelham, 2008, J. Res. Lepid. 40: 318
  - Argynnis adianthe; Barnes & McDunnough, 1917, 8 (missp.)
  - Speyeria adiante; [BOW]: pl. 15, f. 4
  - Speyeria egleis adiaste; [NACL], #4458a
  - Speyeria adiaste adiaste; Pelham, 2008, J. Res. Lepid. 40: 318
- S. a. clemencei (Comstock, 1925) California
  - Argynnis adiaste r. clemencei Comstock, 1925; Bull. South. Calif. Acad. Sci. 24 (3): 67; TL: Atascadero, California
  - Speyeria adiaste clemencei, Butterflies of America [BOA]
  - Speyeria egleis clemencei; [NACL], #4458b; [NHM card]
  - Speyeria adiaste clemencei; Pelham, 2008, J. Res. Lepid. 40: 319
- S. a. atossa (Edwards, 1890) California
  - Argynnis atossa Edwards, 1890; Butts N. Amer. 3: [127], pl. 8, f. 4-5
  - Argynnis atossa f. tejonica Comstock, 1925; Bull. South. Calif. Acad. Sci. 24 (3): 68; TL: Tejon region
    
    = Speyeria egleis atossa ; [NHM card]
    = Speyeria adiaste atossa ; Pelham, 2008, J. Res. Lepid. 40: 319
  - Speyeria adiaste atossa, Butterflies of America [BOA]
  - Argynnis atossa; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Speyeria atossa; [BOW]: pl. 15, f. 8
  - Speyeria egleis atossa; [NACL], #4458c; [NHM card]
  - Speyeria adiaste atossa; Pelham, 2008, J. Res. Lepid. 40: 319

Speyeria atlantis (Edwards, 1862)

ENA. See [maps] Atlantis Fritillary

Argynnis atlantis Edwards, 1862; Proc. Acad. nat. Sci. Philad. 14: 54

Speyeria atlantis, Moth Photographers Group [MPG]
Speyeria atlantis atlantis, Butterflies of America [BOA]
Speyeria atlantis, Butterflies and Moths of North America [BAMONA]

Argynnis atlantis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
Speyeria atlantis; [NACL], #4459; [BOW]: pl. 15, f. 7; [NHM card]; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 319; Dunford, 2009, Insecta Mundi 90: 25; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643

identification uncertain 700x795 (~54Kb) underside USA: FR 9705 nr. US Route 97, (47°18'15N 120°41'24W ±0.5km), Kittitas Co., WA, 14.8.2001, Photo © Markku Savela

identification uncertain 700x976 (~102Kb) underside USA: FR 9705 x US Route 97, (47°18'38N 120°41'48W), Kittitas Co., WA, 14.8.2001, Photo © Markku Savela

Larva on Viola septentrionalis, Viola sororia affinis, V. adunca, V. candensis Dunford, 2009, Insecta Mundi 90: 27

S. a. atlantis Maine, ...
- Argynnis atlantis ab. chemo Scudder, 1889; Butts eastern U.S. Canada 1: 573; TL: Lake Chemo
  - = Speyeria atlantis atlantis ; Pelham, 2008, J. Res. Lepid. 40: 319
  - = Speyeria atlantis ; [NHM card]; Dunford, 2009, Insecta Mundi 90: 25
- Speyeria atlantis atlantis; Pelham, 2008, J. Res. Lepid. 40: 319; Dunford, 2009, Insecta Mundi 90: 27
S. a. canadensis (dos Passos, 1935) Newfoundland, ...
- Argynnis atlantis canadensis dos Passos, 1935; Can. Ent. 67 (4): 85; TL: Doyles Station, Newfoundland
  - = Speyeria atlantis ; Dunford, 2009, Insecta Mundi 90: 25
- Speyeria atlantis canadensis, Butterflies of America [BOA]
- Speyeria atlantis canadensis; [NACL], #4459a; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 319
S. a. hollandi (Chermock & Chermock, 1940) Manitoba, British Columbia
- Argynnis atlantis hollandi Chermock & Chermock, 1940; Can. Ent. 72 (4): 82; TL: Riding Mtns, Manitoba
- Speyeria atlantis hollandi, Butterflies of America [BOA]
- Speyeria atlantis hollandi; [NACL], #4459b; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 319; Dunford, 2009, Insecta Mundi 90: 28
S. a. sorocko Scott, Kondla & Spomer, 1998 Colorado
- Speyeria atlantis sorocko Scott, Kondla & Spomer, 1998; Papilio (N.S) 8: 13; TL: several km NE of Mt. Jude, 9100ft, Clear Creek Co., Colorado
- Speyeria atlantis sorocko, Butterflies of America [BOA]
- Speyeria atlantis sorocko; Pelham, 2008, J. Res. Lepid. 40: 319; Dunford, 2009, Insecta Mundi 90: 28
Larva on Viola sororia affinis, V. scopulorum, V. canadensis Dunford, 2009, Insecta Mundi 90: 29
S. a. pahasapa Spomer, Scott & Kondla, 1998 South Dakota
- Speyeria atlantis pahasapa Spomer, Scott & Kondla, 1998; Papilio (N.S) 8: 13; TL: Deerfield Reservoir, Pennington Co., South Dakota
- Speyeria atlantis pahasapa, Butterflies of America [BOA]
- Speyeria atlantis pahasapa; Pelham, 2008, J. Res. Lepid. 40: 319; Dunford, 2009, Insecta Mundi 90: 29

Speyeria hesperis (Edwards, 1864) Colorado, Utah, Montana. See [maps]
- Argynnis hesperis Edwards, 1864; Proc. ent. Soc. Philad. 2 (4): 502; TL: Rocky Mountains
- Speyeria hesperis, Moth Photographers Group [MPG]
- Speyeria hesperis hesperis, Butterflies of America [BOA]
- Speyeria hesperis, Butterflies and Moths of North America [BAMONA]
- Argynnis hesperis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
- Speyeria hesperis; [BOW]: pl. 16, f. 1; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 319; Dunford, 2009, Insecta Mundi 90: 29; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643
Larva on Viola canadensis var. scopulorum, V. adunca, V. sororia affinis, V. rydbergii, V. adunca bellidifolia, V. nuttallii, V. purpurea Dunford, 2009, Insecta Mundi 90: 30
- S. h. hesperis Colorado
  - Speyeria atlantis hesperis; [NACL], #4459c; [NHM card]
  - Speyeria hesperis hesperis; Pelham, 2008, J. Res. Lepid. 40: 321; Dunford, 2009, Insecta Mundi 90: 29
- S. h. irene (Boisduval, 1869) California, ...
  - Argynnis egleis var. irene Boisduval, 1869; Ann. Soc. ent. Belg. 12: 60
  - Speyeria hesperis irene, Butterflies of America [BOA]
  - Argynnis rupestris irene; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Speyeria atlantis irene; [NACL], #4459n; [NHM card]
  - Speyeria hesperis irene; Pelham, 2008, J. Res. Lepid. 40: 319; Dunford, 2009, Insecta Mundi 90: 34
  Larva on Viola purpurea Dunford, 2009, Insecta Mundi 90: 35
- S. h. nausicaa (Edwards, 1874) Arizona, ...
  - Argynnis nausicaa Edwards, 1874; Trans. amer. ent. Soc. 5 (2): 104; TL: "Rocky Canon, Arizona"
  - Argynnis aphrodite f. arizonensis Elwes, 1889; Trans. ent. Soc. Lond. 1889: 546; TL: Arizona
    
    = Speyeria atlantis nausicaa ; [NHM card]
    = Speyeria hesperis nausicaa ; Pelham, 2008, J. Res. Lepid. 40: 322; Dunford, 2009, Insecta Mundi 90: 42
  - Speyeria hesperis nausicaa, Butterflies of America [BOA]
  - Speyeria atlantis nausicaa, Butterflies of Southeastern Arizon [Bruce Walsh]
  - Speyeria atlantis nausicaa; [NACL], #4459f; [NHM card]
  - Speyeria hesperis nausicaa; Pelham, 2008, J. Res. Lepid. 40: 322; Dunford, 2009, Insecta Mundi 90: 42
- S. h. electa Edwards, 1878 Colorado, Montana, New Mexico
  - Argynnis electa Edwards, 1878; Field & Forest 3: 143
  - Argynnis cornelia Edwards, 1892; Can. Ent. 24 (5): 106
    
    = Speyeria electa ; [NHM card]
    = Speyeria hesperis electa ; Pelham, 2008, J. Res. Lepid. 40: 321; Dunford, 2009, Insecta Mundi 90: 40
  - Argynnis nikias Ehrmann, 1917; Lepidopterist 1 (7): 55; TL: Temez[sic] Springs, New Mexico
    
    = Speyeria hesperis electa ; Pelham, 2008, J. Res. Lepid. 40: 322; Dunford, 2009, Insecta Mundi 90: 41
  - Speyeria hesperis electa, Butterflies of America [BOA]
  - Speyeria atlantis nikias; [NACL], #4459d; [NHM card]
  - Argynnis cornelia; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
  - Argynnis electa; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
  - Speyeria electa; [NHM card]
  - Speyeria atlantis electa; [NACL], #4459o
  - Speyeria hesperis electa; Pelham, 2008, J. Res. Lepid. 40: 321; Dunford, 2009, Insecta Mundi 90: 40
  Larva on Viola sororia affinis Dunford, 2009, Insecta Mundi 90: 41
- S. h. chitone (Edwards, 1879) Utah, Arizona
  - Argynnis chitone Edwards, 1879; Can. Ent. 11 (5): 82; TL: Southern Utah and Arizona
  - Speyeria hesperis chitone, Butterflies of America [BOA]
  - Argynnis chitone; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
  - Speyeria atlantis chitone; [NACL], #4459h; [NHM card]
  - Speyeria hesperis chitone; Pelham, 2008, J. Res. Lepid. 40: 320; Dunford, 2009, Insecta Mundi 90: 40
- S. h. cottlei (Comstock, 1925) California
  - Argynnis cottlei Comstock, 1925; Bull. South. Calif. Acad. Sci. 24 (3): 64; TL: nr Alturas, Modoc Co., California
    
    = Speyeria atlantis irene ; [NHM card]
  - Speyeria hesperis cottlei, Butterflies of America [BOA]
  - Speyeria hesperis cottlei; Pelham, 2008, J. Res. Lepid. 40: 320; Dunford, 2009, Insecta Mundi 90: 35
- S. h. beani (Barnes & Benjamin, 1926) Alberta
  - Dryas atlantis r. beani Barnes & Benjamin, 1926; Bull. South. Calif. Acad. Sci. 25 (3): 92; TL: Banff, Alberta
  - Speyeria hesperis beani, Butterflies of America [BOA]
  - Speyeria atlantis beani; [NACL], #4459r; [NHM card]
  - Speyeria hesperis beani; Pelham, 2008, J. Res. Lepid. 40: 321; Dunford, 2009, Insecta Mundi 90: 31
- S. h. dodgei (Gunder, 1931) Oregon
  - Argynnis dodgei Gunder, 1931; Bull. South. Calif. Acad. Sci. 30 (2): 46; TL: Diamond Lake, Dougla Co., Oregon
  - Speyeria hesperis dodgei, Butterflies of America [BOA]
  - Speyeria atlantis dodgei; [NACL], #4459m; [NHM card]
  - Speyeria hesperis dodgei; Pelham, 2008, J. Res. Lepid. 40: 320; Dunford, 2009, Insecta Mundi 90: 37
  Larva on Violla bellidifolia Dunford, 2009, Insecta Mundi 90: 38
- S. h. lurana dos Passos & Grey, 1945 South Dakota
  - Speyeria atlantis lurana dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 8; TL: Harney Peak, Black Hills, SD
  - Speyeria hesperis lurana, Butterflies of America [BOA]
  - Speyeria atlantis lurana; [NACL], #4459p; [NHM card]
  - Speyeria hesperis lurana; Pelham, 2008, J. Res. Lepid. 40: 321; Dunford, 2009, Insecta Mundi 90: 34
- S. h. tetonia dos Passos & Grey, 1945 Wyoming, Utah
  - Speyeria atlantis wasatchia dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 8; TL: Payson Canyon, Payson, Utah
    
    = Speyeria hesperis tetonia ; Pelham, 2008, J. Res. Lepid. 40: 320
  - Speyeria atlantis tetonia dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 9; TL: Teton Mts., WY
  - Speyeria hesperis tetonia, Butterflies of America [BOA]
  - Speyeria atlantis wasatchia; [NACL], #4459i; [NHM card]
  - Speyeria hesperis wasatchia; Dunford, 2009, Insecta Mundi 90: 39
  - Speyeria atlantis tetonia; [NACL], #4459k; [NHM card]
  - Speyeria hesperis tetonia; Pelham, 2008, J. Res. Lepid. 40: 320; Dunford, 2009, Insecta Mundi 90: 39
- S. h. viola dos Passos & Grey, 1945 Idaho
  - Speyeria atlantis viola dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 10; TL: Trail Creek, Sawtooth Mts., ID, 7400ft
  - Speyeria hesperis viola, Butterflies of America [BOA]
  - Speyeria atlantis viola; [NACL], #4459l; [NHM card]
  - Speyeria hesperis viola; Pelham, 2008, J. Res. Lepid. 40: 320; Dunford, 2009, Insecta Mundi 90: 37
- S. h. hutchinsi dos Passos & Grey, 1947 Montana
  - Speyeria atlantis hutchinsi dos Passos & Grey, 1947; Amer. Mus. Novit. 1370: 20; TL: Twp 2N Range 1W, Jefferson Co., Montana
    
    = Speyeria hesperis beani ; Dunford, 2009, Insecta Mundi 90: 31
  - Speyeria atlantis beani tr.f. hutchinsi Gunder, 1932; Can. Ent. 64 (12): 280; TL: Twp 2N Range 1W, Jefferson Co., Montana
    
    = Speyeria hesperis hutchinsi ; Pelham, 2008, J. Res. Lepid. 40: 320
  - Speyeria hesperis hutchinsi, Butterflies of America [BOA]
  - Speyeria atlantis hutchinsi; [NACL], #4459q; [NHM card]
  - Speyeria hesperis hutchinsi; Pelham, 2008, J. Res. Lepid. 40: 320
- S. h. dorothea Moeck, 1947 New Mexico
  - Speyeria atlantis dorothea Moeck, 1947; Ent. News 58 (3): 73; TL: Sandia Peak, Sandia Mtns, New Mexico
  - Speyeria hesperis dorothea, Butterflies of America [BOA]
  - Speyeria atlantis dorothea; [NACL], #4459e; [NHM card]
  - Speyeria hesperis dorothea; Pelham, 2008, J. Res. Lepid. 40: 322; Dunford, 2009, Insecta Mundi 90: 43
  Larva on Viola sororia affinis Dunford, 2009, Insecta Mundi 90: 43
- S. h. schellbachi Garth, 1949 Arizona, ...
  - Speyeria atlantis schellbachi Garth, 1949; Bull. South. Calif. Acad. Sci. 48 (1). 1-2: 1; TL: Neal Springs, North Rim, Grand Canyon National Park, Coconino Co., Arizona
  - Speyeria hesperis schellbachi, Butterflies of America [BOA]
  - shellbachi; dos Passos, 1964, (missp.)
  - Speyeria atlantis schellbachi; [NACL], #4459g; [NHM card]
  - Speyeria hesperis schellbachi; Pelham, 2008, J. Res. Lepid. 40: 320; Dunford, 2009, Insecta Mundi 90: 41
- S. h. greyi Moeck, 1950 Nevada
  - Speyeria atlantis greyi Moeck, 1950; Ent. News 61 (3): 61; TL: Lamoille Canyon, Ruby Mtns, Elko Co., Nevada
  - Speyeria hesperis greyi, Butterflies of America [BOA]
  - Speyeria atlantis greyi; [NACL], #4459j; [NHM card]
  - Speyeria hesperis greyi; Pelham, 2008, J. Res. Lepid. 40: 320; Dunford, 2009, Insecta Mundi 90: 33
- S. h. helena dos Passos & Grey, 1957 Alberta, British Columbia, Manitoba
  - Argynnis lais Edwards, 1883; Can. Ent. 15 (11): 209 (preocc. Argynnis lais Scudder, 1875); TL: "Red Deer River"
    
    = Speyeria atlantis helena ; [NHM card]
    = Speyeria hesperis helena ; Pelham, 2008, J. Res. Lepid. 40: 321; Dunford, 2009, Insecta Mundi 90: 31
  - Argynnis lais tr.f. dennisi Gunder, 1927; Can. Ent. 59 (12): 287; TL: Beulah, Manitoba
    
    = Speyeria hesperis helena ; Pelham, 2008, J. Res. Lepid. 40: 321; Dunford, 2009, Insecta Mundi 90: 31
  - Speyeria atlantis dennisi dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: 20; TL: Beulah, Manitoba
    
    = Speyeria hesperis helena ; Pelham, 2008, J. Res. Lepid. 40: 321; Dunford, 2009, Insecta Mundi 90: 31
  - Speyeria (Speyeria) atlantis helena dos Passos & Grey, 1957; J. N.Y. Ent. Soc. 63: 95 (repl. Argynnis lais Edwards, 1883)
  - Speyeria atlantis dennisi; [NACL], #4459t
  - Speyeria electa dennisi; [NHM card]
  - Argynnis lais; Dyar, 1903, Bull. U.S. nat. Mus. 52: 13
  - Speyeria lais; [BOW]: pl. 16, f. 3
  - Speyeria atlantis lais; [NACL], #4459s
  - Speyeria atlantis helena; [NHM card]
  - Speyeria hesperis helena; Pelham, 2008, J. Res. Lepid. 40: 321; Dunford, 2009, Insecta Mundi 90: 31
- S. h. ratonensis Scott, 1981 New Mexico
  - Speyeria atlantis ratonensis Scott, 1981; Papilio (n.s.) 1: 1; TL: Raton Mesa, Colfax Co., New Mexico
  - Speyeria hesperis ratonensis, Butterflies of America [BOA]
  - Speyeria hesperis ratonensis; Pelham, 2008, J. Res. Lepid. 40: 321; Dunford, 2009, Insecta Mundi 90: 33
- S. h. elko Austin, 1983 Nevada (Elko Co.)
  - Speyeria atlantis elko Austin, 1983; J. Lep. Soc. 37 (3): 244, f. 1-2; TL: Nevada, Elko Co., Owybee River Valley, Wild Horse Creek Camping, ca. 10mi S of Mountain City
  - Speyeria hesperis elko, Butterflies of America [BOA]
  - Speyeria hesperis elko; Pelham, 2008, J. Res. Lepid. 40: 320; Dunford, 2009, Insecta Mundi 90: 38
- S. h. capitatensis Holland, 1988 New Mexico
  - Speyeria atlantis capitanensis Holland, 1988; Bull. Allyn Mus. 115: 2; TL: Padilla Point, crest of Capitan Ridge, Capitan Mts, Lincoln Co., NM
  - Speyeria hesperis capitatensis; Pelham, 2008, J. Res. Lepid. 40: 322; Dunford, 2009, Insecta Mundi 90: 43
- S. h. hanseni Emmel, Emmel & Mattoon, 1998 California
  - Speyeria atlantis hanseni Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (12): 152; TL: California, Tehama Co., Anthony Peak on Covelo Rd
  - Speyeria hesperis hanseni, Butterflies of America [BOA]
  - Speyeria hesperis hanseni; Pelham, 2008, J. Res. Lepid. 40: 319; Dunford, 2009, Insecta Mundi 90: 36
- S. h. brico Kondla, Scott & Spomer, 1998 British Columbia
  - Speyeria hesperis brico Kondla, Scott & Spomer, 1998; Papilio (N.S) 8: 14; TL: Castle Creek FS Rd Km 23.5, Cariboo Mts, British Columbia
  - Speyeria hesperis brico, Butterflies of America [BOA]
  - Speyeria hesperis brico; Pelham, 2008, J. Res. Lepid. 40: 320; Dunford, 2009, Insecta Mundi 90: 32

Speyeria hydaspe (Boisduval, 1869)

W.USA. See [maps] Hydaspe Fritillary

Argynnis hydaspe Boisduval, 1869; Ann. Soc. ent. Belg. 12: 60; TL: California

Speyeria hydaspe, Moth Photographers Group [MPG]
Speyeria hydaspe, Butterflies of America [BOA]
Speyeria hydaspe, Butterflies and Moths of North America [BAMONA]

Speyeria hydaspe; [NACL], #4460; [EBW]; [NHM card]; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 322; Dunford, 2009, Insecta Mundi 90: 44; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643

800x828 (~114Kb) underside USA: Taneum Creek, Wenatchee National Forest, (47°06'24N 120°54'44W), Kittitas Co., WA, 29.7.1999, Photo © Markku Savela	identification uncertain 800x758 (~67Kb) upperside USA: FR9705, Table Mtn, Kittitas Co., WA, 30.7.1999, Photo © Markku Savela	500x673 (~59Kb) male USA: End of FR 6520 /Johnson Ridge, Mt. Baker Snoqualmie Nat. Forest (47°47'48N 121°15'45W), Snohomish Co., WA, 7.8.2001, Photo © Markku Savela
500x582 (~46Kb) underside USA: End of FR 6520 /Johnson Ridge, Mt. Baker Snoqualmie Nat. Forest (47°47'48N 121°15'45W), Snohomish Co., WA, 7.8.2001, Photo © Markku Savela

Larva on Viola adunca, V. glabella, V. nuttallii, V. orbiculata, V. purpurea, V. sheltonii Dunford, 2009, Insecta Mundi 90: 45

S. h. hydaspe California
- Argynnis monticola var. purpurascens H. Edwards, 1877; Proc. Calif. Acad. Sci. 7 (12): 170; TL: Siskiyou Co.
  - = Speyeria hydaspe hydaspe ; Pelham, 2008, J. Res. Lepid. 40: 322
- Argynnis hydaspe ab. caliginosa Comstock, 1925; Bull. South. Calif. Acad. Sci. 24 (3): 66; TL: Modoc Co., California
  - = Speyeria hydaspe purpurascens ; [NHM card]
  - = Speyeria hydaspe hydaspe ; Pelham, 2008, J. Res. Lepid. 40: 322
- Argynnis monticola purpurascens; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
- Speyeria hydaspe purpurascens; [NACL], #4460b
- Speyeria hydaspe hydaspe; Pelham, 2008, J. Res. Lepid. 40: 322
S. h. rhodope (Edwards, 1874) Vancouver, British Columbia
- Argynnis rhodope Edwards, 1874; Trans. amer. ent. Soc. 5 (1): 13; TL: Cariboo District, British Columbia
- Argynnis sakuntala Skinner, 1911; Ent. News 22 (3): 108; TL: Kaslo, B. C.
  - = Speyeria hydaspe rhodope ; Pelham, 2008, J. Res. Lepid. 40: 323
- Argynnis skinneri Holland, 1931; The Butterfly Book (1931 edn): 95, pl. 56, f. 4 (repl. Argynnis sakuntala Skinner, 1911); TL: Kaslo, B. C.
  - = Speyeria hydaspe sakuntala ; [NHM card]
  - = Speyeria hydaspe rhodope ; Pelham, 2008, J. Res. Lepid. 40: 323
- Speyeria hydaspe conquista dos Passos & Grey, 1945; Amer. Mus. Novit. 1297: (1-17); TL: Little Tesuque Canyon, nr Sante Fe, New Mexico, 8000ft
  - = Speyeria hydaspe rhodope ; Pelham, 2008, J. Res. Lepid. 40: 323
- Speyeria hydaspe rhodope, Butterflies of America [BOA]
- Speyeria hydaspe conquista; [NACL], #4460f; [NHM card]
- Speyeria hydaspe sakuntala; [NACL], #4460e; [NHM card]
- Argynnis rhodope; Dyar, 1903, Bull. U.S. nat. Mus. 52: 14
- Speyeria hydaspe rhodope; [NACL], #4460d; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 323
S. h. viridicornis (Comstock, 1925) California
- Argynnis hydaspe f. viridicornis Comstock, 1925; Bull. South. Calif. Acad. Sci. 24 (3): 63; TL: Greenhorn Mtns, California
- Speyeria hydaspe viridicornis, Butterflies of America [BOA]
- Speyeria hydaspe viridicornis; [NACL], #4460a; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 322
S. h. minor dos Passos & Grey, 1947 British Columbia
- Speyeria (Speyeria) hydaspe minor dos Passos & Grey, 1947; Amer. Mus. Novit. 1370: 21; TL: Mt. McLean, B. C.
- Argynnis rhodope tr.f. gregsoni Gunder, 1932; Can. Ent. 64 (12): 281; TL: Mt Washington, Forbidden Plateau, V.I., B.C
  - = Speyeria hydaspe rhodope ; [NHM card]
  - = Speyeria hydaspe minor ; Pelham, 2008, J. Res. Lepid. 40: 323
- Argynnis hydaspe rhodope f. alt. minor McDunnough, 1927; Can. ent. 59: 155; TL: Mt. McLean, B. C.
  - = Speyeria hydaspe minor ; Pelham, 2008, J. Res. Lepid. 40: 323
- Speyeria hydaspe minor, Butterflies of America [BOA]
- Speyeria hydaspe minor; [NACL], #4460c; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 323
S. h. tehama Emmel, Emmel & Mattoon, 1998 California
- Speyeria hydaspe tehama Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (12): 146; TL: California, Tehama Co., Covelo Rd, at Govenrment Flat, 5900'
- Speyeria hydaspe tehama, Butterflies of America [BOA]
- Speyeria hydaspe tehama; Pelham, 2008, J. Res. Lepid. 40: 322
S. h. davisi Emmel, Emmel & Mattoon, 1998 California
- Speyeria hydaspe davisi Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (12): 147; TL: California, Humboldt Co., McClellan Mtn peat bog
- Speyeria hydaspe davisi, Butterflies of America [BOA]
- Speyeria hydaspe davisi; Pelham, 2008, J. Res. Lepid. 40: 323

Speyeria mormonia (Boisduval, 1869) California, Nevada. See [maps] Mormon Fritillary
- Argynnis mormonia Boisduval, 1869; Ann. Soc. ent. Belg. 12: 58
- Speyeria mormonia, Lepiforum [LEPIFORUM]
- Speyeria mormonia, Moth Photographers Group [MPG]
- Speyeria mormonia, Butterflies of America [BOA]
- Speyeria mormonia, Butterflies and Moths of North America [BAMONA]
- Speyeria mormonia; [NACL], #4461; [EBW]; [BOW]: pl. 16, f. 4; [NHM card]; [Opler]; Pelham, 2008, J. Res. Lepid. 40: 323; Dunford, 2009, Insecta Mundi 90: 45; de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017, Syst. Ent. 42 (4): 643
1200x885 (~136Kb) upperside USA: Washington, Skagit Co., Sauk Mountain Trail Head & Trail, 15.8.2012, Photo © Markku Savela

Larva on Viola nuttallii, V. palustris, V. adunca, Viola adunca var. bellidifolia, V. sororia Dunford, 2009, Insecta Mundi 90:# 46
- S. m. mormonia Oregon?, California
  - Argynnis astarte Edwards, 1862; Proc. Ent. Soc. Philad. 1 (7): 221 (preocc. by Doubleday, 1847)
    
    = Speyeria mormonia arge ; [NHM card]
    = Speyeria mormonia mormonia ; Pelham, 2008, J. Res. Lepid. 40: 325
  - Argynnis montivaga Behr, 1864; Proc. Ent. Soc. Philad. 3 (3): 435 (preocc. Argynnis montivaga Behr, 1863)
    
    = Speyeria mormonia arge ; [NHM card]
    = Speyeria mormonia mormonia ; Pelham, 2008, J. Res. Lepid. 40: 325
  - Argynnis arge Strecker, 1878; Synon. Cat. Macroplep.: 114
    
    = Speyeria mormonia mormonia ; Pelham, 2008, J. Res. Lepid. 40: 325
  - Argynnis eurynome arge; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Speyeria mormonia arge; [NACL], #4461f; [NHM card]
  - Speyeria mormonia mormonia; Pelham, 2008, J. Res. Lepid. 40: 325
- S. m. bischoffii (Edwards, 1870) Alaska
  - Argynnis bischoffii Edwards, 1870; Trans. amer. ent. Soc. 3: 189; TL: Anchorage, Alaska
  - Speyeria mormonia bischoffii, Butterflies of America [BOA]
  - Argynnis eurynome bischoffi; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Speyeria mormonia bischoffi; [NHM card]
  - Speyeria mormonia bischoffii; [NACL], #4461a; Grey, 1989, J. Lep. Soc. 43 (1): 6; Pelham, 2008, J. Res. Lepid. 40: 323
- S. m. eurynome (Edwards, 1872) Wyoming, Montana, Alberta, Pacific Coast, ...
  - Argynnis eurynome Edwards, 1872; Trans. amer. ent. Soc. 4: 66
  - Argynnis clio Edwards, 1874; Trans. amer. ent. Soc. 5 (2): 106
    
    = Speyeria mormonia eurynome ; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 324
  - Argynnis eurynome ab. eris Igel, 1922; (preocc. by Meigen, 1828)
    
    = Speyeria mormonia eurynome ; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 324
  - Argynnis mormonia tr.f. benjamini Gunder, 1927; Can. Ent. 59 (12): 287; TL: Arangie, Idaho
    
    = Speyeria mormonia eurynome ; Pelham, 2008, J. Res. Lepid. 40: 324
  - Argynnis eurynome tr.f. brucei Gunder, 1927; Can. Ent. 59 (12): 287
    
    = Speyeria mormonia eurynome ; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 325
  - Argynnis eurynome r. clio f. gunderi Field, 1936; J. Ent. Zool. 28 (2): 23 (preocc. by Comstock, 1925); TL: Broadwater Co., Montana
    
    = Speyeria mormonia eurynome ; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 325
  - Speyeria mormonia eurynome ab. igeli dos Passos & Grey, 1947; Amer. Mus. Novit. 1370: 23 (repl. Argnynnis eurynome ab. eris Igel, 1922)
    
    = Speyeria mormonia eurynome ; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 325
  - Speyeria mormonia eurynome ab. fieldi dos Passos & Grey, 1947; Amer. Mus. Novit. 1370: 23 (repl. Argynnis eurynome clio trans.f. gunderi Field, 1936)
    
    = Speyeria mormonia eurynome ; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 325
  - Speyeria mormonia eurynome, Butterflies of America [BOA]
  - eurynomes; Edwards, 1873, (missp.)
  - Speyeria mormonia benjamini; [NHM card]
  - Argynnis eurynome clio; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Argynnis eurynome; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Speyeria eurynome; [BOW]: pl. 15, f. 18
  - Speyeria mormonia eurynome; [NACL], #4461h; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 324
- S. m. opis (Edwards, 1874) British Columbia
  - Argynnis opis Edwards, 1874; ; TL: Bald Mt., Cariboo Dist., British Columbia
  - Argynnis bischoffi opis f. jesmondensis McDunnough, 1940; Can. ent. 72 (2): 25; TL: Jesmond, B. C.
    
    = Speyeria mormonia opis ; Grey, 1989, J. Lep. Soc. 43 (1): 6; Pelham, 2008, J. Res. Lepid. 40: 324
  - Speyeria mormonia jesmondensis dos Passos & Grey, 1947; Amer. Mus. Novit. 1370: 22; TL: Jesmond, B. C.
    
    = Speyeria mormonia opis ; Pelham, 2008, J. Res. Lepid. 40: 324
  - Speyeria mormonia opis, Butterflies of America [BOA]
  - Speyeria mormonia jesmondensis; [NACL], #4461c; [NHM card]
  - Speyeria mormonia opis; [NACL], #4461b; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 324
- S. m. artonis (Edwards, 1881) Nevada, ...
  - Argynnis artonis Edwards, 1881; Trans. Amer. Ent. Soc. 9: 1
  - Argynnis eurynome artonis f. rubyensis Gunder, 1932; Can. Ent. 64 (12): 283; TL: Franklin Slough, nr Wright's Ranch, Ruby Valley, Elko Co., Nevada
    
    = Speyeria mormonia artonis ; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 325
  - Speyeria mormonia artonis, Butterflies of America [BOA]
  - Argynnis eurynome artonis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Speyeria mormonia artonis; [NACL], #4461g; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 325
- S. m. erinna (Edwards, 1883) Washington, Oregon
  - Argynnis eurynome var. erinna Edwards, 1883; Can. Ent. 15 (2): 33; TL: Spokane Falls
  - Argynnis erinna ab. cunninghami Owen, 1893; Ent. News 4 (7): 246; TL: Klamath Co., Oregon
    
    = Speyeria mormonia erinna ; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 324
  - Speyeria mormonia erinna, Butterflies of America [BOA]
  - Argynnis eurynome cunninghami; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Argynnis eurynome erinna; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15
  - Speyeria mormonia erinna; [NACL], #4461e; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 324
  Larva on Viola palustris, Viola adunca var. bellidifolia Hammond & Dornfeld, 1983, J. Lep. Soc. 37 (2): 120
- S. m. washingtonia (Barnes & McDunnough, 1913) Washington
  - Argynnis bischoffi[sic] washingtonia Barnes & McDunnough, 1913; Contr. nat. Hist. Lep. N. Am. 2 (3): 95, pl. 1, f. 5; TL: Mt Ra[i]nier, 7000ft, [Pierce Co.], Wash[ington]
  - Argynnis eurynome washingtonia f. ranierensis Gunder, 1932; Can. Ent. 64 (12): 282; TL: Paradise Valley, Mount Ra[i]nier, Pierce Co., Washington
    
    = Speyeria mormonia washingtonia ; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 324
  - Speyeria mormonia washingtonia, Butterflies of America [BOA]
  - ranierensis; Guppy & Kondla, 2002, J. Lep. Soc. 56 (3): 181
  - Speyeria rainierensis[sic]; [NACL], #4461
  - Speyeria mormonia washingtonia; [NACL], #4461d; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 324
- S. m. luski (Barnes & McDunnough, 1913) Arizona
  - Argynnis eurynome luski Barnes & McDunnough, 1913; Contr. nat. Hist. Lep. N. Am. 2 (3): 95; TL: White Mts, Arizona
  - Speyeria mormonia luski, Butterflies of America [BOA]
  - Speyeria mormonia luski; [NACL], #4461i; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 326
- S. m. obsidiana Emmel, Emmel & Mattoon, 1998 California
  - Speyeria mormonia obsidiana Emmel, Emmel & Mattoon, 1998; Syst. W. N.Am. Butts. (12): 145; TL: California, Mono Co., Sawmill Meadow, 9200', Glass Mtn
  - Speyeria mormonia obsidiana, Butterflies of America [BOA]
  - Speyeria mormonia obsidiana; Pelham, 2008, J. Res. Lepid. 40: 325

Unknown or unplaced taxa

?albomaculata (Rebel)
- = Speyeria aglaja ; [NHM card]

22.4.2019 (19)

References:

[BAMONA] Opler, Pavulaan, Stanford & Pogue
Butterflies and Moths of North America; Mountain Prairie Information Node
[BCA] electronic
BIOLOGIA CENTRALI-AMERICANA
[BOA] Interactive Listing of American Butterflies
Listado Interactivo de la Mariposas Americanas; Butterflies of America
[Bruce Walsh]
Bruce Walsh; e-mail: Bruce Walsh; The Moths of Southeastern Arizona; Butterflies of Southeastern Arizona
[CiNii]
http://ci.nii.ac.jp/
[D. Smirnov]
Russia, Moscow; e-mail: D. Smirnov
[FLMNH] Florida Museum of Natural History
Bulletin of the Allyn Museum
[GALEPS] Moths and Butterflies of Georgia and the Southeastern United States
James K. Adams; Dept. of Natural Science, Dalton State College, Dalton, GA ; http://www.daltonstate.edu/galeps/index.htm
[Heiner Ziegler]
Switzerland / Schweiz; www.pieris.ch
[JLS]
Journal of the Lepidopterists' Society; the original J. Lep. Soc. articles
[Kimmo Silvonen] Espoo, Finland
Kimmo's Lep Site
[LAMAS-2005] Gerardo Lamas
Entomofauna 26 (6): 57-100; A Bibliography of the Zoological Publications of Hans Fruhstorfer (1886* - 1922+)
[LEPIFORUM]
Lepiforum; Bestimmung von Schmetterlingen (Lepidoptera) und ihren Präimaginalstadien
[MBE] Moths and Butteflies of Europe
Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[MPG]
North American Moth Photographers Group
[Mario Maier]
Europäische Schmetterlinge; e-mail: Mario Maier
[Michael Borsch]
Geesthacht, Germany; contact through; e-mail: Francisco Welter-Schultes
[Michel Tarrier]
Michel R. Tarrier; Malaga, Spain; http://www.tarrier.org
[NBC] Netherlands Centre for Biodiversity Naturalis
NBC naturalis
[NHM] Natural History Museum
The Global Lepidoptera Names Index; Butterflies and Moths of the World; Generic Names and their Type-species
[Opler] Opler & Warren, 2003 (April)
Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.
[Pertti Pakkanen]
e-mail: Pertti Pakkanen; Butterfly and moth observations from Finland
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)
[Simon Coombes] 1 Park Street, Plymouth, PL3 4BL, UK
01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[Tero Piirainen] Tampere, Finland
e-mail: Tero Piirainen
[Vitaly Charny]
Birmingham, Alabama, USA
[Wahlberg]
Niklas Wahlberg; The NSG's voucher specimen database; Nymphalidae Systematics Group
[ZOBODAT] Oberösterreichische Landesmuseum
OÖ. Landesmuseum; ZOBODAT - Literatur
[maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.
[²] This may require parentheses or not. I don't have the necessary information for this taxon.

Some related literature:

Acerbi, 1802
Travels through Sweden, Finland and Lapland to North Cape, in the years 1798 and 1799 Travels North Cape 2 : 3-380, pl. 1-3
Austin, 1983
A new subspecies of Speyeria atlantis from the great basin of Nevada J. Lep. Soc. 37 (3) : 244-248
Austin, 1998
New subspecies of Nymphalidae (Lepidoptera) from Nevada and Arizona Syst. W. N. Am. Butts. (46) : 573-586
Barnes, 1897
Some new species and varieties of Lepidoptera from the Western U. S. Can. Ent. 29 (2) : 39-42
Barnes & Benjamin, 1926
Notes on diurnal Lepidoptera, with addition and corrections to the recent "List of Diurnal Lepidoptera" Bull. South. Calif. Acad. Sci. 25 (3) : 88-98
Barnes & McDunnough, 1913
New North American Lepidoptera with notes on described species Contr. nat. Hist. Lep. N. Am. 2 (3) : 93-165, pl. 1-9, I-IX
Behr, 1862
[Our California Argynnides] Proc. Cal. Acad. Sci. 2 : 172-177
Behr, 1863
On Californian Lepidoptera - No. III Proc. Calif. Acad. Sci. 3 (1) : 84-93
Behr, 1864
Notes on the Argynnides of California Proc. Ent. Soc. Philad. 3 (3) : 434-436
Boisduval, 1852
Lépidoptères de la Californie Ann. Soc. ent. Fr. (2) 10 (2) : 275-324
Boisduval, 1869
Lépidoptères de la Californie Ann. Soc. ent. Belg. 12 : 5-28,37-94
Bramson, 1890
Die Tagfalter (Rhopalocera) Europas und des Caucasus Tagfalter Eur. Caucasus : 1-148
Brown, 1983
The type of Argynnis apacheana Skinner J. Lep. Soc. 37 (1) : 79-80
Chermock & Frechin, 1947
A new Speyeria from Washington Pan-Pacific Ent. 23 (3) : 111-112
[MRS]; Chou, 1994
Monografia Rhopalocerorum Sinensium 1-2: 1-854
Chou, Yuan, Yin, Zhang & Chen, 2002
New species, new subspecies, and new records of butterflies from China (VI) Entomotaxonomia 24 (1): 52-68
Clark, 1932
The butterflies of the District of Columbia and vicinity Bull. U.S. nat. Mus. 157 : 1-337
Cockerell, 1900
A New Variety of Argynnis Nitocris Ent. News 11 (10) : 622
Cockerell, 1909
Notes on Butterflies Ent. Rec. J. Var. 21 (7/8) : 186
Comstock, 1925
Studies in Pacific Coast Lepidoptera Bull. South. Calif. Acad. Sci. 24 (1): 3-4, (2): 35-37, pl. 2, (3): 62-68
Cramer, [1775]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 1 (1-7): 1-132, pl. 1-84 (1775), (8): 133-155, pl. 85-96 (1776)
Cramer, [1777]
Uitlandsche Kapellen (Papillons exotiques) Uitl. Kapellen 2 (9-16): 1-152, pl. 97-192 (1777)
de Nicéville, 1889
On new and little-known Butterflies from the Indian Region, with Revision of the Genus Plesioneura of Felder and of Authors J. Bombay nat. Hist. Soc. 4 (3) : 163-194, pl. A-B
dos Passos, 1935
Some butterflies of southern Newfoundland with descriptions of new subspecies Can. Ent. 67 (4) : 82-88
dos Passos & Grey, 1942
Two new North American subspecies of Argynnis, with some revisional notes Amer. Mus. Novit. 1214: 6pp, 1fig
dos Passos & Grey, 1945
A new species and some new subspecies of Speyeria Amer. Mus. Novit. 1297: 17pp, 30fgs
dos Passos & Grey, 1947
Systematic catalogue of Speyeria with designations of types and fixations of type localities Amer. Mus. Novit. 1370: 30pp
dos Passos & Grey, 1957
A new name for Argynnis lais Edwards J. N.Y. Ent. Soc. 63: 95-96
Drury, [1773]
Illustrations of natural history; wherein are exhibited ... Illust. Nat. Hist. Exot. Insects 1 : 1-130, pl. 1-50 (1770), (1773)
Dunford, 2009
Taxonomic overview of the greater fritillary genus Speyeria Scudder and the atlantis - hesperis species complexes, with species accounts, type images, and relevant literature (Lepidoptera: Nymphalidae) Insecta Mundi 90 : 1-74
Dyar, 1903
A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723
Edwards, 1862
Descriptions of certain Species of Diurnal Lepidoptera found within the limits of the United States and of British America. - No. [1]-3 Proc. Acad. nat. Sci. Philad. 13: 160-164 (1861), 14: 54-58, 221-226 (1862)
Edwards, 1862
Descriptions of certain species of Diurnal Lepidoptera found within the United States, figured in Doubleday's Genera but undescribed Proc. Ent. Soc. Philad. 1 (7) : 221-224
Edwards, 1864
Descriptions of certain species of Diurnal Lepidoptera, found within the limits of the United States and British America. (No. 1-3) Proc. ent. Soc. Philad. 2 (1): 14-22, pl. 1, 11 (1863), (2): 78-82 (1863), (4): 501-507 (1864)
Behr in Edwards, 1864; Edwards, 1864
Notes on the Argynnides of California Proc. ent. Soc. Philad. 3 : 434-436
Edwards, 1869
The Butterflies of North America. Vol. I Butts. N. Amer. 1: 218pp, 50pls
Edwards, 1890
The Butterflies of North America. Vol. III Butts N. Amer. 3: 431pp, 51pls
Edwards, 1869
Descriptions of new species of diurnal Lepidoptera found within the United States Trans. amer. ent. Soc. 2 (4) : 369-376
Edwards, 1870
Descriptions of new species of diurnal Lepidoptera found within the United States Trans. amer. ent. Soc. 3 (1) : 10-22
Edwards, 1870
Descriptions of new North American diurnal Lepidoptera Trans. amer. ent. Soc. 3 : 189-196
Edwards, 1872
Descriptions of new species of diurnal Lepidoptera found within the United States Trans. amer. ent. Soc. 4 : 61-70
Edwards, 1874
Descriptions of new species of diurnal Lepidoptera found in North America Trans. amer. ent. Soc. 5 (1): 13-19, (2): 103-111
Edwards, 1876
Description of new species of diurnal Lepidoptera found within the United States and British North America Trans. amer. ent. Soc. 5 : 202-208
Edwards, [1877]
New species of diurnal Lepidoptera Trans. amer. ent. Soc. 5 : 289-292
Edwards, 1877
Descriptions of new species of diurnal Lepidoptera found in North America Field & Forest 3 (5): 86-89, : 101-105
Edwards, 1878
Descriptions of new species of North American Lepidoptera Field & Forest 3 : 115-119, 142-144
Edwards, 1879
Descriptions of new species of butterflies collected by Mr. H. K. Morrison in Nevada, 1878; also, Remarks on some errors of synonymy and arrangement Can. Ent. 11 (3): 49-56, : 79-80
Edwards, 1879
Descriptions of new species of North American butterflies; also, Notes upon certain species Can. Ent. 11 (5) : 81-89
Edwards, 1881
Descriptions of new species of diurnal Lepidoptera found within the United States Trans. Amer. Ent. Soc. 9 : 1-8
Edwards, 1883
Descriptions of new species of diurnal Lepidoptera found in British America and the United States Can. Ent. 15 (2) : 32-36
Edwards, 1883
Descriptions of new species of North American butterflies Can. Ent. 15 (11) : 209-211
Edwards, 1886
Description of new species of butterflies found in the United States Can. Ent. 18 (4) : 61-65
Edwards, 1892
Miscellaneous notes on butterflies, larvae, etc. Can. Ent. 24 (3): 49-56, (5): 105-111
H. Edwards, 1877
Pacific Coast Lepidoptera. - No. 22. Notes on some Diurnal Lepidoptera, with description of new varieties Proc. Calif. Acad. Sci. 7 (12) : 163-173
H. Edwards, 1877
Pacific Coast Lepidoptera. No. 26. - Description of two new forms of Diurnals Proc. Cal. Acad. Sc.: 1 (Dec. 17)
Ehrmann, 1917
Some New North American Butterflies Lepidopterist 1 (7) : 54-56
Eisner, 1942
Einige ergebnisse der Sichtung der Gattungen Melitaea und Argynnis im Rijksmuseum van Natuurlijke Historie, Leiden Zool. Meded. Leiden 24 (4) : 116-124
Elwes, 1889
A revision of he genus Argynnis Trans. ent. Soc. Lond. 1889 : 535-575
Evans, 1912
A list of Indian butterflies J. Bombay nat. Hist. Soc. 21 (2): 553-584, (3): 969-1008
Evans, 1924
The identification of Indian Butterflies (5-8) J. Bombay nat. Hist. Soc. 30 (1): 72-96, pl. 21-24, (2): 322-351, pl. 26-27, (3): 610-639, pl. 28, (4): 726-776, pl. 29
Fabricius, 1775
Systema Entomologiae, sistens Insectorum Classes, Ordines, Genera, Species, Adiectis Synonymis, Locis, Descriptionibus, Observationibus Syst. Ent. : 1-832
Fabricius, 1787
Mantissa insectorum sistens species nuper detectas adiectis synonymis, observationibus, descriptionibus, emendationibus Mantissa Insectorum 2 : 1-382
Fawcett, 1904
On some New and Little-known Butterflies, mainly from high elevations in the N.E. Himalayas Proc. zool. Soc. Lond. 2 (1) : 131-141, pl. 9
Fisher, 1858
Description of a New Species of ARGYNNIS Proc. Acad. Nat. Sci. Philad. 10 : 179-180
Fruhstorfer, 1903
Neue Nymphaliden aus Ostasien Dt. ent. Z. Iris 16 (1) : 22-42, pl. 1, f. 5-6 (early October)
Fruhstorfer, 1906
Rhopaloceren-Aberrationen Ent. Zs. 20 (3) : 17-18, f. 1-5 (15 April)
Fruhstorfer, 1907
Neue Argynnis aus der aglaja-Gruppe Int. ent. Zs. 1 (34) : 256-257 (16 November)
Fruhstorfer, 1908
Eine neue Argynnis-Rasse aus Spanien Ent. Zs. 22 (39) : 161 (26 December)
Fruhstorfer, 1912
6. Familie: Nymphalidae in Seitz, Gross-Schmett. Erde 9 : 453-456 (31 January, 1912), : 457-488 (18 March, 1912), : 489-496 (25 May, 1912), : 489-496 (10 May, 1912), : 497-504 (15 April, 1912), : 505-512 (10 May, 1912), : 513-528 (11 April, 1912), : 529-552 (20 April, 1912), : 553-592 (15 May, 1912), : 593-616 (25 September, 1912), : 617-632 (15 October, 1912), : 633-640 (25 October, 1912), : 641-656 (10 January, 1913), : 657-664 (12 January, 1913), : 665-680 (13 January, 1913), : 681-688 (25 January, 1913), : 689-696 (16 June, 1913), : 697-704 (5 August, 1913), : 705-712 (20 January, 1914), : 713-720 (20 November, 1913), : 721-744 (9 March, 1914), : 745-751 (30 July, 1915), : 753-760 (28 March, 1914), : 761-766 (14 April, 1914)
Godman & Salvin, [1901]
Biologia Centrali-Americana. Rhopalocera. (1887-1901) Biol. centr.-amer., Lep. Rhop. 2: 1-782, 3: pl. 1-112
Grey, 1989
Sundry Argynnine concepts revisited (Nymphalidae) J. Lep. Soc. 43 (1): 1-10
Grum-Grshimailo, 1890
Le Pamir et sa faune lépidoptérologique in Romanoff, Mém. Lép. 4 : 1-576, pl. A,1-21
Gunder, 1924
Several New Aberrant Lepidoptera (Rhopalocera) from California Ent. News 35 (5) : 153-158
Gunder, 1925
Several New Varieties of and Aberrant Lepidoptera (Rhopalocera) from California Ent. News 36 : 1-9, pl. 1
Gunder, 1929
New butterflies and sundry notes (Lepidoptera, Rhopalocera) Bull. Brooklyn ent. Soc. 24 (5) : 325-332, pl. 30-31
Gunder, 1931
Some new Butterflies (Lepid., Rhopalocera) Bull. South. Calif. Acad. Sci. 30 (2) : 45-48
Gunder, 1932
New Rhopalocera Can. Ent. 64 (12) : 276-284
Gunder, 1933
Additional New Rhopalocera (Lepidoptera) Can. Ent. 65 (8) : 171-173
Guppy & Kondla, 2002
The correct spellings and type localities of butterflies named from Mount Rainier, Washington, USA J. Lep. Soc. 56 (3): 180-131
Hammond & Dornfeld, 1983
A new subspecies of Speyeria egleis from the pumice region of central Oregon J. Lep. Soc. 37 (2): 115-120
Hemming, 1933
Holarctic butterflies: miscellaneous notes on nomenclature Entomologist 66 (12) : 275-279
Hemming, 1942
On the correct name of the species commonly known as Argynnis aglaja (Linnaeus, 1758) (Lep. Nymphalidae) and matters incidental hereto Proc. R. ent. Soc. Lond. (B) 11 (11) : 155-160
[NACL]; Hodges, 1983
Check List of the Lepidoptera of America North of Mexico Check List Lep. Am. N of Mexico
Holland, 1900
A Description of a Variety of Argynnis Nitocris from Chihuahua, Mexico Ent. News 11 (1) : 332-333
Holland, 1931
Notes on some American butterflies mainly relating to classification and nomenclature. Part 2 & 3 Ann. Carnegie Mus. 20 (1): 39-55 (1930), (2): 255-264 (1931)
Holland, 1988
A new subspecies of Speyeria atlantis (Nymphalidae) from south-central New Mexico Bull. Allyn Mus. 115: 1-9
Holland, 2010
A new subspecies of Argynnis nokomis from Sacramento Mountains of New Mexico J. Res. Lepid. 42: 79-80
Holmgren, 1888
Aberrationer till Argynnis aglaja L och adippe L. Ent. Tidskr. 9 : 103-106
Jachontov, 1909
[Notice sur les Lèpidoptères Rhopalocères du Caucase] Revue Russe Ent. 8 : 282-292
Janson, 1877
Notes on Japanese Rhopalocera with the description of new species Cistula ent. 2 : 153-160
Kardakoff, 1928
Zur Kenntnis der Lepidopteren des Ussuri-Gebietes Ent. Mitt. 17 (4): 261-273, (8): 414-425
Kershaw, 1952
Two New Aberrations of Argynnis aglaia Linn. Ent. Rec. J. Var. 64 : 69-70, pl. 3
Korb & Bolshakov, 2011
[A catalogue of butterflies (Lepidoptera: Papilioformes) of the former USSR. Second edition, reformatted and updated] (in Russian) Eversmannia Suppl. 2 : 1-123
Kudrna, 1974
An annotated List of Japanese Butterflies Atalanta 5 : 92-120
Kuwayama, 1921
On some Aberrations of the Japanese Butterflies Dobuts. Z. 33 : 332-337
[NL4A]; Lamas, 2004
Checklist: Part 4A, Hesperioidea - Papilionoidea Atlas of Neotropical Lepidoptera
Lampa, 1885
Förteckning öfver Skandinaviens och Finlands Macrolepidoptera Ent. Tidskr. 6 (1) : 1-137
Lang & Han, 2009; Lang in Lang & Han, 2009
Study on some nymphalid butterflies from China Atalanta 40 (3/4) : 493-500
Lang in Lang & Wang, 2010
Study on some nymphalid butterflies from China - 2 Atalanta 41 : 221-228
[RFEL]; Leley, 2016
Annotated catalogue of the insects of Russian Far East. Volume II. Lepidoptera Cat. ins. Russian Far East. 2 : 1-812
Lempke, 1956
Catalogus der Nederlandse Macrolepidoptera (Vierde Supplement) Tijdschr. Ent. 99 (4) : 155-216
[BOW]; Lewis, 1974
Butterflies of the World
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Linnaeus, 1767
Systema Naturae per Regna tria Naturae, secundum Classes, Ordines, .... Editio Duocecima Reformata. Tom. 1. Part II. Syst. Nat. (Edn 12) 1 (2) : 533-1327, : Errata
Matsumura, 1908
Die Nymphaliden Japans Ent. Zs. 22 (39) : 157-158
Matsumura, 1911
Erster Beitrag zur Insekten Fauna von Sachalin J. Coll. Agr. Tohoku Imp. Univ. Sapporo 4 (1) : 1-145, pl. 1-2
Matsumura, 1928
New Butterflies especially from the Kuriles Insecta Matsumurana 2 (4) : 191-201
Matsumura, 1929
Some new butterflies from Korea received from Mr. T. Takamuku Insecta Matsumurana 3 (4) : 152-156
McCorkle & Hammond, 1988
Biology of Speyeria zerene hippolyta (Nymphalidae) in marine-modified environment J. Lep. Soc. 42 (3): 184-195
McDunnough, 1924
Some new Canadian argynnid races (Lepid.) Can. ent. 56 (2) : 42-43
McDunnough, 1927
The Lepidoptera of the Seton Lake Region, British Columbia Can. ent. 59 : 152-162,193-199,207-214,239-246,266-277
McDunnough, 1935
A new race of Argynnis cybele from Nova Scotia Can. ent. 67: 18-19
McDunnough, 1940
The argynnids of the Cariboo region of British Columbia (Lepid.) Can. ent. 72 (2) : 23-25
Ménétriés, 1832
Catalogue raisonne des objects de zoologie recuellis dans un voyage au Caucase et jusqu'aux frontières de la Perse Cat. Rais. : 1-271
Moeck, 1947
A New Subspecies of Speyeria atlantis (Edwards) from New Mexico (Lepidoptera: Nymphalidae) Ent. News 58 (3) : 73-75
Moeck, 1950
A New Subspecies of Speyeria atlantis (Edwards) from Nevada (Lepidoptera: Nymphalidae) Ent. News 61 (3) : 61-64
Moeck, 1957
Geographic variability in Speyeria. Comments, records and description of a new subspecies Milwaukee Ent. Soc., Special Paper: -
Moore, [1875]
Descriptions of new Asiatic Lepidoptera Proc. zool. Soc. Lond. 1874 (4) : 565-579, pl. 66-67
de Moya, Savage, Tenney, Bao, Wahlberg & Hill, 2017
Interrelationships and diversification of Argynnis Fabricius and Speyeria Scudder butterflies Syst. Ent. 42 (4) : 635-649
Nakahara, 1926
Some Saghalien Butterflies: Notes and Descriptions Ins. Inscit. Mens. 14 : 45-48
Oberthür, 1909
Notes pour servir à établir la Faune Francaise et Algeriénne des Lépidoptères Étud. Lépid. Comp. 3 : 101-400, 405-415, pl. 11,16-20,23-26
Owen, 1893
Peculiar form of Argynnis erinna Edw Ent. News 4 (7) : 246
Pelham, 2008
A catalogue of the butterflies of the United States and Canada J. Res. Lepid. 40 : 1-652
Peschke, 1934
Neubeschreibungen und Ergänzungen der palaearktischen Lepidopterenfauna I. Int. ent. Zs. 28 : 430-432
Pflümer, 1879
Ein Beitrag zur Schmetterlingskunde Stettin Ent. Ztg 40 : 157-161
Reakirt, 1866
Coloradian butteflies Proc. Ent. Soc. Philad. 6 (1) : 122-151
Reiff, 1910
Argynnis cybele Fabr. Forma bartschi f. nov. Psyche 17 (6) : 252
Reuss, 1926
Systemischer uberlich der Dryadinae T. Rss mit einigen Neubeschreibungen Dt. ent. Zs. 1926 (1) : 65-70
Reuss, 1926
Über Funktion der Sexualarmaturen bei Lepidopteren (Rhop.) und die resultierende Weiterentwicklung meines versuchten natürlichen Systems der Dryadinae T. R. Dt. ent. Zs. 1926 (5) : 431-440
Riel, 1911
Argynnis aglaja L. aberr. hortensia Donzel Ann. Soc. Linn. Lyon (n.s.) 58 : 187-188
Rothschild, 1933
On a collection of Lepidoptera from Spanish Morocco Novit. Zool. 38 : 315-330
Röber, 1896
Neue Schmetterlinge aus dem cilicischen Taurus Ent. Nachr. 22 (6) : 81-84
de Sagarra, 1925
Anotacions a la Lepidopterologica ibèrica Butll. Inst. catal. Hist. nat. (2) 5 (1): 45-49, (9): 270-274
de Sagarra, 1926
Anotacions a la lepidopterologia Ibérica IV (1) Butll. Inst. catal. Hist. nat. (2) 6 (6-7) : 128-139
de Sagarra, 1930
Anotacions a la lepidopterologia Ibérica V (2). Formes noves de lepidòpters ibérics Butll. Inst. catal. Hist. nat. (2) 10 (7) : 110-118
Schuster von Forstner, 1928
Völlig schwarzer Perlmutterfalter Lepid. Rdsch. 2 (16) : 157-158
Scott, 1981
New Papilionoidea and Hesperioidea from North America Papilio (n.s.) 1 : 1-12
Kondla, Scott & Spomer in Scott, Kondla & Spomer, 1998; Scott, Kondla & Spomer, 1998; Spomer, Scott & Kondla in Scott, Kondla & Spomer, 1998
Speyeria hesperis and Speyeria atlantis are distinct species Papilio (N.S) 8 : 1-31
Scudder, 1872
A systematic Revision of some of the American Butterflies; with brief notes on those known to occur in Essex County, Mass. 4th Ann. Rep. Peabody Acad. Sci. (1871) : 24-83,errata
Scudder, 1875
Synonymic list of the butterflies of North Amercia, north of Mexico. (1) Nymphales Bull. Buffalo Soc. nat. Sci. 2 : 233-269
Scudder, 1889
The Butterflies of the Eastern United States and Canada, with special reference to New England Butts eastern U.S. Canada 1: 1-766, 2: 767-1774, 3: 1775-1958, pl. 1-89
[SPRK]; Seppänen, 1970
Suomen suurperhostoukkien ravintokasvit Animalia Fennica 14
Skinner, 1897
Notes on Rhopalocera, with descriptions of new species and varieties Can. Ent. 29 (6) : 154-156
Skinner, 1911
A new Argynnis and a new Parnassius (Lep.) Ent. News 22 (3) : 108
Skinner, 1917
New species of Lepidoptera Ent. News 28 (7) : 328-329
Skinner, 1918
Argynnis apacheana, a New Name (Lepid.) Ent. News 29 (2) : 67
Skinner, 1919
A New Species of Argynnis from Utah (Lepid., Rhop.) Ent. News 30 (8) : 216
Slaby, 1949
Faunistické paberky z Cech. (Lep.) - Faunistiques Lépidoptèlogogiques de la Boheme Acta Soc. ent. Cal. 45 (1-2) : 65-72
[EBW]; Smart, 1976
The Illustrated Encyclopedia of the Butterfly World Illust. Encyp. Butt. World: -
Staudinger in Staudinger & Rebel, 1901
Catalog der Lepidopteren des palaearktischen Faunengebiets. I. Famil. Papilionidae - Hepialidae Cat. Lep. palaearct. Faunengeb. 1 : 1-411
Stichel, 1901
Berichtigung und Ergänzung zu meinem Artikel über bemerkenswerthe Schmetterlings-Varietäten etc. Berl. ent. Z. 46 : 21-24
Strand, 1901
Beitrag zur Schmetterlingsfauna Norwegens Nyt Mag. Naturv. 39 (1) : 25-72
Strand, 1912
Ueber einige Nebenformen europäischer Großschmetterlinge Ent. Zs. 25 (49): 253-254, (50): 257-258
Strecker, 1878
Butterflies and Moths of North America. Complete Synonymical Catalogue of Macrolepidoptera ... Synon. Cat. Macroplep.: - : 1-281
[BMAT]; Tennent, 1996
The Butterflies of Morocco, Algeria and Tunisia : 1-217
Tutt, 1896
British Butterflies, being a popular hand-book for young stutends and collectors Brit. Butts : 1-476
[BRU2]; Tuzov, Bogdanov, Churkin, Devyatkin, Dantchenko, Murzin, Samodurov, Zhdanko, 2000
Guide to the Butterflies of Russia and adjacent territories: Libytheidae, Danaidae, Nymphalidae, Riodinidae, Lycaenidae Butts. Russia adj. terr. 2 : 1-580
Verity, 1914
Contributo allo studio della variazione nei Lepidotteri Boll. Soc. ent. ital. 45 : 203-238, pl. 1
Verity, 1919
Seasonal polymorphism and races of some European Grypocera and Rhopalocera Ent. Rec. J. Var. 31 : 26-31,43-48,87-89,121-129,178-184,193-201
Verity, 1922
Seasonal polymorphism and races of some European Grypocera and Rhopalocera Ent. Rec. J. Var. 34 : 12-15, 68-73, 89-93, 124-142
Verity, 1935
Notes on the geographical variations of Argynnis aglaja, L. Entomologist 68 : 189-191, : 200-204
Wileman, 1911
Aberrations of three Japanese butterflies Entomologist 44 : 55-56
Williams, 2002
Recognition of western populations of Speyeria idalia (Nymphalidae) as new subspecies J. Lep. Soc. 55 (4) : 144-149
Wolcott, 1916
Description of two Hitherto Undescribed Aberrations, the one of Argynnis alcestis, the of the Chlorippe celtis (Lep.) Ent. News, 27 (3) : 97-99, pl. 4
Wright, 1905
The Butterflies of the West Coast of the United States Butts. West Coast US : 3-257
Yakovlev, 2012
Checklist of Butterflies (Papilionoidea) of the Mongolian Altai Mountains, including descriptions of new taxa Nota lepid. 35 (1) : 51-96

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Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.