-
Boloria selene (Denis & Schiffermüller, 1775)
. See [maps]
Niittyhopeatäplä
Small Pearl-bordered Fritillary
Silver-bordered Fritillary
Silver Meadow Fritillary
Brunfläckig pärlemorfjäril
Ängspärlefjäril
Braunfleckiger Perlmutterfalter
Le Petit Collier Argenté
Perlada castaña
Dostojka selene
- Papilio selene Denis & Schiffermüller, 1775; Ank. syst. Schmett. Wienergegend: 321; TL: Austria
- Papilio euphrosyne Bergsträsser, 1779; Nomen. Ins. 3: 42, f. 1-2 [?]
- Papilio euphrasia Lewin, 1795; Pap. Gr. Britain: 30, pl. 13, f. 3-4
- Papilio marphisa Herbst, 1800; Naturs. Schmett. 10: 105, pl. 278, f. 8-9
- Papilio rinaldus Herbst, 1800; Naturs. Schmett. 10: 108, pl. 271, f. 1-2, 4
- Papilio silene Haworth, 1803; Lepid. Britannica (1): 34
- Argynnis selenia Freyer, 1850; Neuere Beitr. Schmett. 6: pl. 493, f. 2
- Brenthis selene var. intermedia Spångberg, 1876; Lepid. Anteckn.: 26
- transversa (Tutt, 1896) (Brenthis); Brit. Butts: 295
- Argynnis selene f. (gen. II) angustipennis Fuchs, 1899; Jb. Nassau Ver. Nat. 52: 123
- Argynnis selene ab. flavopunctata Strand, 1900; Arch. Nat. Christian 21 (5): 20
- Argynnis selene ab. flavopunctata Strand, 1900; Ent. Nachr. 26 (15 & 16): 226
- Argynnis selene ab. pallida Spuler, 1901; Schmett. Eur. 1: 26
- Argynnis selene ab. nigromarginata Lambillion, 1906; Rev. Mens. Soc. ent. namur. 6 (10): 48
- Argynnis selene ab. gerda Schultz, 1908; Ent. Zs. 22 (9): 38
- Brenthis selene var. castiliana Lowe, 1910; Ent. Rec. J. Var. 22: 23; TL: La Granja
- Argynnis selene lucetia Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 19; TL: Japan
- interligata (Cabeau, 1919) (Argynnis); Rev. mens. Soc. ent. Namur. 1919: 49
- halflantsi (Cabeau, 1922) (Argynnis); Rev. mens. Soc. ent. Namur. 1922: 17
- consuta (Cabeau, 1922) (Argynnis); Rev. mens. Soc. ent. Namur. 1922: 50
- extenuata (Cabeau, 1922) (Argynnis); Rev. mens. Soc. ent. Namur. 1922: 50
- Argynnis selene ab. semipallida Nessling, 1925; Notulae Ent. 5 (3): 97; TL: Finland, Vetil, Röyrinki
- furoshonis (Matsumura, 1929) (Argynnis); Ill. comm. Ins. Japan 1: 16 (repl. sachalinensis Matsumura)
- Clossiana selene f. h. futura Reuss, 1929; Int. ent. Z. 22: 433
- medionigrans (Cabeau, 1930) (Argynnis); Lambillionea 30: 54
- vanescens (Cabeau, 1930) (Argynnis); Lambillionea 30: 54
- obsoleta (Curtis, 1933) (Argynnis); Trans. ent. Soc. S. Engl. 8: 105
- Boloria selene (II) gen. postgentilina Verity, 1933; Bull. Soc. ent. Fr. 38 (8): 116
- Boloria selene r. padimaxima Verity, 1933; Bull. Soc. ent. Fr. 38 (8): 117
- Brenthis selene ab. striata Lempke, 1936; Tijdschr. Ent. 79: 269; TL: Groningen
- Argynnis selene ab. e-nigrum Foltin, 1938; Zs. Öst. EntVer. 23: 125; TL: Kefermarkt
- pseudoselene (Rocci, 1940) (Argynnis); Mem. Soc. ent. ital. 19: 51
- Argynnis selene ab. novo plesiensis Jesátko, 1944; Acta Soc. ent. bohem. 41: 146
- Clossiana selene f. loc. hyperborea Lingonblad, 1947; Notulae Ent. 26 (3-4): 69; TL: Finland, Kilpisjärvi
- Clossiana selene padimaxima f. postpadimaxima Verity, 1950; Le Farfalle diurn. d'Italia 4: 234
- = Boloria selene postgentilinea ; [NHM card]
- Argynnis selene ab. fasciata Schnaider, 1950; Polskie Pismo. Ent. 19 (3-4): 241
- Clossiana selene f. flavescens Lempke, 1956; Tijdschr. Ent. 99 (4): 158
- Clossiana selene f. debrunneata Lempke, 1956; Tijdschr. Ent. 99 (4): 159
- Clossiana selene f. infraclara Lempke, 1956; Tijdschr. Ent. 99 (4): 159
- Clossiana selene f. infradefasciata Lempke, 1956; Tijdschr. Ent. 99 (4): 160
- Clossiana selene f. semicadmeis Lempke, 1956; Tijdschr. Ent. 99 (4): 160
- Brenthis selene var. intermedia; Spångberg, 1878, Bihang K. svenska VetensAkad. Handl. 5: 8
- Boloria selene angustipennis; [NHM card]
- Boloria selene castiliana; [NHM card]
- Boloria selene debrunneata; [NHM card]
- Argynnis selene ab. flavopunctata; Strand, 1912, Ent. Zs. 25 (49): 254
- Boloria selene flavescens; [NHM card]
- Boloria selene flavopunctata; [NHM card]
- Boloria selene futura; [NHM card]
- Boloria selene hyperborea; [NHM card]
- Boloria selene infraclara; [NHM card]
- Boloria selene infradefasciata; [NHM card]
- Boloria selene lucetia; [NHM card]
- Boloria selene marphisa; [NHM card]
- Boloria selene postgentilinea[sic, recte postgentilina]; [NHM card]
- Boloria selene pseudoselene; [NHM card]
- Clossiana selene semicademis[sic, recte semicadmeis]; [NHM card]
- Clossiana selene; [NACL], #4464; [BOW]: pl. 2, f. 2-3; [EBW]; [BARU, #188]; [BRU2]: 45, pl. 32, f. 1-6; Yakovlev, 2012, Nota lepid. 35 (1): 90; [RFEL]
- Boloria (Clossiana) selene; Pelham, 2008, J. Res. Lepid. 40: 298
- Boloria selene; [NHM card]; [BNA]: 318, #239, pl. 3, 5, 24, 26; [BOC], 281; [Opler]; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
![[selene-1u.jpg]](_selene-1u.jpg)
450x697 (~67Kb) upperside
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![[selene-1v.jpg]](_selene-1v.jpg)
458x713 (~67Kb) underside
|
![[selene-2u.jpg]](_selene-2u.jpg)
425x374 (~35Kb) upperside
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![[selene-2v.jpg]](_selene-2v.jpg)
401x353 (~34Kb) underside
|
![[selene-3u.jpg]](_selene-3u.jpg)
500x391 (~75Kb) upperside Haldon woods, Devon, UK 21/05/95, Photo © Simon Coombes
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![[selene-3v.jpg]](_selene-3v.jpg)
500x389 (~57Kb) underside Haldon woods, Devon, UK 21/05/95, Photo © Simon Coombes
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![[selene-4v.jpg]](_selene-4v.jpg)
837x1028 (~120Kb) underside Finland, Kuhmalahti, Mattilankylä, Setälänlahti, 23.6.2005, Photo © Harri Arkkio
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![[selene-4vx.jpg]](_selene-4vx.jpg)
1371x992 (~199Kb) underside Finland, Kuhmalahti, Mattilankylä, Setälänlahti, 23.6.2005, Photo © Harri Arkkio
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![[selene-5fu.jpg]](_selene-5fu.jpg)
852x639 (~123Kb) upperside female Russia, Moscow area (36°25'E, 56°00'N), 14.6.2009, Photo © D. Smirnov
|
![[selene-5mu.jpg]](_selene-5mu.jpg)
711x624 (~103Kb) upperside male Russia, Moscow area, 14.6.2010 (36°25'E, 56°00'N), Photo © D. Smirnov
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![[selene-5v.jpg]](_selene-5v.jpg)
824x689 (~114Kb) underside Russia, Moscow area, 10.6.2010 (36°25'E, 56°00'N), Photo © D. Smirnov
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![[selene-6mu.jpg]](_selene-6mu.jpg)
918x694 (~122Kb) upperside male Russia, Moscow area, 3.7.2010 (36°25'E, 56°00'N), Photo © D. Smirnov
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![[selene-6mv.jpg]](_selene-6mv.jpg)
726x732 (~106Kb) underside male Russia, Moscow area, 3.7.2010 (36°25'E, 56°00'N), Photo © D. Smirnov
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![[selene-7u.jpg]](_selene-7u.jpg)
700x700 (~78Kb) upperside A grassy open larch forest on the Yabaganskii Pass (1400 m above sea level) through the Seminskii mountain range, the northern Altai Mts., West Siberia, Russia. 11th July 1999, Photo © Oleg Kosterin
|
![[selene-8u.jpg]](_selene-8u.jpg)
916x545 (~97Kb) upperside Russia, Moscow area, 15.6.2011 (36°25'E, 56°00'N), Photo © D. Smirnov
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Larva on
Viola ,
V. palustris,
V. canina,
V. riviniana [SPRK],
Fragaria ,
Vaccinium uliginosum [BRU2], 46,
Viola nephrophylla,
V. glabella [BNA], 318
-
B. s. selene EU, W.Siberia, N.Kazakhstan, Altai, Sayan, Transbaikalia
-
B. s. myrina (Cramer, [1777])
- Papilio myrina Cramer, [1777]; Uitl. Kapellen 2 (9-16): 141, pl. 189, f. B, C
- Papilio myrinus Herbst, 1798
- Argynnis myrissa Godart, [1824]; Encyclopédie Méthodique 9 (1): 253
- Brenthis selene marilandica Clark, 1941; J. Wash. Acad. Sci. 31 (9): 384; TL: Beltsville, Maryland
- Brenthis myrina ab. nubes Scudder, 1889; Butts eastern U.S. Canada 1: 595; TL: Sanbornton, New Hampshire
- Brenthis myrina tr.f. nivea Gunder, 1928; Can. Ent. 60 (7): 163 (albino); TL: Staten Island, New York
- Brenthis selene ab. gunderana Forbes, 1960; Cornell Univ. Agric. Exp. Stn. Mem. 371: 150
- Clossiana myrina nivea; [NHM card]
- Boloria selene marilandica; Klots, 1951, 89; dos Passos, 1964, 89
- Brenthis myrina ab. nubes; McDunnough, 1938, 16
- Argynnis myrina; Scudder, 1863, 166; Scudder, 1868, 379; Edwards, 1876, 161
- Brenthis myrina; Dyar, 1900, 487; Cary, 1901, 307; Dyar, 1903, Bull. U.S. nat. Mus. 52: 15; Cary, 1906, 439; Elrod, 1906, 69; Franzen, 1914, 366; McDunnough, 1928, 274; Holland, 1931, The Butterfly Book (1931 edn): 105; Clark, 1932, 102; dos Passos & Grey, 1934, 190; McDunnough, 1938, 15; Davenport & Dethier, 1938, 158; Leussler, 1938, 216; Rysgaard, 1939, 193; Comstock, 1940, 53; Brooks, 1942, 34
- Boloria myrina; dos Passos & Grey, 1945, 4; [Joseph Belicek]
- Brenthis selene myrina; Clark, 1941, 384
- Clossiana selene myrina; [NACL], #4464a; [BNA], #239
- Clossiana myrina; [NHM card]
- Boloria selene myrina; Klots, 1951, 89; dos Passos, 1964, 89; Shepard, 1975, in Howe, 246; Kohler, 1977, J. Lep. Soc. 31 (4): 245; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 298
-
B. s. thalia (Esper, 1789) Arctic Europe, N.Siberia
-
B. s. chibiana (Matsumura, 1927) Sakhalin, Kunashir
-
B. s. tollandensis (Barnes & Benjamin, 1925) Rocky Mountains (Colorado, Idaho, Montana, New Mexico, Utah, Wyoming), British Columbia
- Brenthis myrina tollandensis Barnes & Benjamin, 1925; Ent. News 36: 44; TL: Tolland, Gilpin Co., Colorado
- Brenthis myrina tollandensis; Holland, 1928, 37; Klots, 1937, 328; McDunnough, 1938, 16
- Brenthis myrina var. tollandensis; Holland, 1931, The Butterfly Book (1931 edn): 106
- Brenthis selene tollandensis; Clark, 1941, 384
- Boloria myrina tollandensis; Brown, 1954, 64
- Clossiana myrina tollandensis; [NHM card]
- Boloria selene tollandensis; Remington, 1952, 67; Brown, 1957, 64; Albright, 1960, 158; Newcomer & Rogers, 1963, 171; dos Passos, 1964, 89; Shepard, 1975, in Howe, 246; Kohler, 1977, J. Lep. Soc. 31 (4): 253
- Clossiana selene tollandensis; [NACL], #4464d
- Boloria selene tollandensis; Pelham, 2008, J. Res. Lepid. 40: 299
-
B. s. atrocostalis (Huard, 1927) Nova Scotia, Quebec, Saskatchewan, Manitoba, Maine, Vermont, New Hampshire, New York, Michinag, Wisconsin, Minnesota, North Dakota
- Argynnis atrocostalis Huard, 1927; Nat. Can. 54 (6): 131; TL: Chicoutini, Quebec
- Brenthis myrina ab. ♂ serratimarginata Gunder, 1926; Ent. News 37 (1): 7; TL: Vernon, British Columbia
- Brenthis myrina ab. jenningsae Holland, 1928; Ann. Carnegie Mus. 19 (1): 36; TL: Jellicoe, on Thunder Bay, Ontario
- Brenthis myrina var. jenningsae Holland, 1931; The Butterfly Book (1931 edn): 105, pl. LVI, f. 9 (type)
- Brenthis myrina atrocostalis ab. jenningsae; McDunnough, 1932, 269; McDunnough, 1938, 16
- Boloria selene atrocostalis ab. jenningsae; dos Passos, 1964, 89
- Brenthis myrina atrocostalis; McDunnough, 1930, 107; McDunnough, 1938, 16; dos Passos & Grey, 1934, 190
- Clossiana selene atrocostalis; [NACL], #4464f
- Boloria selene atrocostalis; Clark, 1941, 384; Klots, 1951, 89; dos Passos, 1954, 89; Shepard, 1975, in Howe, 246; Kohler, 1977, J. Lep. Soc. 31 (4): 258, f. 5, 8; [BNA], 318; [NHM card]; Pelham, 2008, J. Res. Lepid. 40: 299
-
B. s. nebraskensis (Holland, 1928) E.Nebraska - Ohio
- Brenthis myrina var. nebraskensis; Holland, 1931, The Butterfly Book (1931 edn): 105, pl. LV, f. 8 (type)
- Brenthis myrina r. nebraskensis; Leussler, 1938, 216
- Brenthis myrina nebraskensis; McDunnough, 1938, 16
- Brenthis selene nebraskensis; Clark, 1941, 384
- Clossiana selene nebraskensis; [NACL], #4464b
- Boloria selene nebrascensis; Forbes, 1960, 160 (missp.)
- Boloria selene nebraskensis; Klots, 1951, 89; Brown, 1957, 64; dos Passos, 1964, 89; Shepard, 1975, in Howe, 246; Kohler, 1977, J. Lep. Soc. 31 (4): 248, f. 2, 6; [BNA], #239; Pelham, 2008, J. Res. Lepid. 40: 299
-
B. s. albequina (Holland, 1928) Yukon, SW.Alaska
- Brenthis myrina atrocostalis ab. baxteri; McDunnough, 1938, 16
- Boloria selene albequina ab. baxteri; dos Passos, 1964, 89
- Clossiana albequina baxteri; [NHM card]
- Brenthis albequina; Holland, 1931, The Butterfly Book (1931 edn): 107, pl. LV, f. 19-21 (types); McDunnough, 1932, 269
- Clossiana selene albequina; [NACL], #4464e
- Clossiana albequina; [NHM card]
- Boloria selene albequina; dos Passos, 1964, 89; Kohler, 1977, J. Lep. Soc. 31 (4): 255, f. 4; Pelham, 2008, J. Res. Lepid. 40: 299
-
B. s. terraenovae (Holland, 1928) Newfoundland
- Brenthis myrina terrae-novae; dos Passos, 1935, 87; McDunnough, 1938, 16
- Brenthis myrina var. terrae-novae; Holland, 1931, The Butterfly Book (1931 edn): 105, pl. LV, f. 13 (m.type)
- Clossiana myrina terraenovae; [NHM card]
- Brenthis selene terrae-novae; Clark, 1941, 384
- Clossiana selene terraenovae; [NACL]
- Boloria selene terrae-novae; Klots, 1951, 89
- Boloria selene terraenovae; dos Passos, 1964, 89; Shepard, 1975, in Howe, 264; Kohler, 1977, J. Lep. Soc. 31 (4): 262, f. 5; Pelham, 2008, J. Res. Lepid. 40: 299
-
B. s. sugitanii (Seok, 1938) Korea
- Argynnis selene sugitanii Seok, 1938; Zephyrus 7: 241; TL: Korea
-
B. s. sabulocollis (Kohler, 1977) Nebraska, South Dakota, North Dakota, Colorado
- Boloria selene sabulocollis Kohler, 1977; J. Lep. Soc. 31 (4): 252, f. 3, 6-7; TL: Smith Lake, Sheridan Co., Nebraska
-
Boloria perryi (Butler, 1882)
S.Ussuri, N.Korea, Amur?. See [maps]
-
Boloria jerdoni (Lang, 1868)
W.Himalayas, Chitral - Kashmir. See [maps]
Jerdons Silverspot
- Clossiana jerdoni chitralensis; [NHM card]
- Clossiana jerdoni; [BOW]: pl. 194, f. 13; [NHM card]
-
Boloria erubescens (Staudinger, 1901)
-. See [maps]
-
-
B. e. tienschanica (Wagner, 1913) Tian-Shan
- Argynynnis hegemone tienschanica Wagner, 1913; Ent. Mitt. 2 (4): 114, f. 9; TL: Juldus
- = Clossiana hegemone hegemone ; [BRU2], 46
![[tienschanica-1v.jpg]](_tienschanica-1v.jpg)
400x533 (~49Kb) underside An alpine meadow on the right bank terrace of the fourth tributary of the Sargir River (the Usek River basin), altitude 3200 m, 47 km NEN of Zharkent, 21 km NEN of village Sarybel', the mountain massif Tyshkantau, the Southern Dzhungarian Alatau mountain chain (NE part of Tien Shan), Zharkent District, Taldy-Kurgan Province, SE Kazakhstan. 27th of July 1994, Photo © Oleg Kosterin
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-
B. e. chotana (Bang-Haas, 1915) Pakistan, ...
- Argynnis hegemone var. chotana Bang-Haas, 1915; Dt. ent. Z. Iris 29 (2/3): 100; TL: Kun-Lun Mts., China
- Clossiana franciscana Verhulst, 1999; Lambillionea 99 (2): 213; TL: Pakistan, Gilgit
- Clossia hegemone ali Churkin, 2002; Helios 3: 162; TL: Alai
-
B. e. haberhaueri Hemming, 1933 Alatau, Margelan
- Argynnis hegemone Staudinger, 1881; Stettin ent. Ztg 42 (7-9): 292 (preocc. Argynnis hegemone Godart, 1819); TL: "Ala Tau" [Dzhungarsky Alatau Mts., Kazakhstan]
- = Clossiana erubescens haberhaueri ; [NHM card]
- Argynnis hegemone Alphéraky, 1881; Horae Soc. ent. Ross. 16 (3-4): 407, pl. 15, f. 16-17 (Staudinger i.l.); TL: Kuldja?
- = Clossiana erubescens haberhaueri ; [NHM card]
- Boloria haberhaueri Hemming, 1933; Entomologist 66 (12): 275 (repl. Argynnis hegemone Staudinger, 1881)
-
B. e. houri Wyatt, 1961 W.Tian-Shan, Ghissar-Darvaz, Pamirs-Alai
- Boloria hegemone houri Wyatt, 1961; J. Lep. Soc. 15 (1): 8; TL: Bala Quran, Anjuman Valley, Hindu Kush Mts., 14300ft
-
B. e. eximia (Churkin, 2002) Fergansky Mts
- Clossiana hegemone eximia Churkin, 2002; Helios 3: 164; TL: Fergansky Mts, 15km E Kalmak-Krchin v., Kugart R., 2000-2200m
-
B. e. tuzovi (Churkin, 2002) Talassky Alatau
- Clossiana hegemone tuzovi Churkin, 2002; Helios 3: 165; TL: Talassky Alatau, Alabel Pass, 3300m
-
B. e. ancilla (Churkin, 2004) Tadjikistan
- Clossiana hegemone ancilla Churkin, 2004; Helios 5: 132; TL: Tadjikistan, Peter the Great Range, 23km SE Tadzhikobad, Ganishou v.
-
B. e. panterai Korb, 2005
- Boloria erubescens panterai Korb, 2005; A catalogue of butterflies of the ex-USSR: 86
-
Boloria selenis (Eversmann, 1837)
-. See [maps]
Larva on
Viola [BRU2], 44
-
B. s. selenis S.Urals, Volga Valley
- Clossiana selenis selenis
-
B. s. sibirica (Erschoff, 1870) Amur, NE.Siberia
![[sibirica-1v.jpg]](_sibirica-1v.jpg)
532x326 (~51Kb) underside A southern steppefied slope of the mountain on the Chuya River right bank between the villages Kurai and Chagan-Uzun, Altai Mts., West SIberia, Russia. 7th July 1988, Photo © Oleg Kosterin
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-
B. s. onorensis (Matsumura, 1925) Sakhalin
- Boloria selenis onorensis; [NHM card]
- Clossiana selenis onorensis; [BRU2], 44
-
Boloria oscarus (Eversmann, 1844)
-. See [maps]
![[oscarus-1mv.jpg]](_oscarus-1mv.jpg)
488x400 (~38Kb) underside male A bushy meadow on the second right bank terrace of the Koen River at the Volchikha Rivulet just upstream of its fall into the Koen River near the village Nizhniy Koen, Iskitim District, Novosibirsk Province, West Siberia, Russia. 18th June, 1989, Photo © Oleg Kosterin
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Larva on
Viola [BARU]
-
B. o. oscarus S.Siberia, E.Amur
- Clossiana oscarus oscarus
-
B. o. australis (Graeser, 1888) S.Amur, Ussuri, Sakhalin
-
B. o. oscaroides (Ménétriés, 1859) E.Yakutia
-
Boloria euphrosyne (Linnaeus, 1758)
. See [maps]
Pursuhopeatälä
Pearl-bordered Fritillary
Prydlig pärlemorfjäril
Skogspärlefjäril
Silberstrich
Grand Collier Argenté
Perlada rojiza
Dostojka eufrozyna
- Papilio euphrosyne Linnaeus, 1758; Syst. Nat. (Edn 10) 1: 481, no. 142
- Papilio euphrosyne Linnaeus, 1761; Fauna Suecica (Edn 2): 282; TL: Sweden
- Papilio argenticollis Retzius, 1783; Gen. Spec. Ins.: 31
- Brenthis morrisii Reakirt, 1866; Proc. Acad. nat. Sci. Philad. 18 (3): 246; TL: California [error]
- obsoleta (Tutt, 1896) (Brenthis); Brit. Butts: 300
- pauca (Tutt, 1896) (Brenthis); Brit. Butts: 300
- Argynnis euphrosyne v. apennina Staudinger, 1901; Cat. Lep. palaearct. Faunengeb. 1: 35
- Argynnis euphrosyne ab. melanotica Spuler, 1901; Schmett. Eur. 1: 27
- Argynnis euphrosyne ab. pallida Spuler, 1901; Schmett. Eur. 1: 27
- Argynnis euphrosyne ab. radiata Spuler, 1901; Schmett. Eur. 1: 27
- Argynnis euphrosyne ab. tatrica Aigner-Abafi, 1906; Ann. hist.-nat. Mus. Hung. 4 (2): 502; TL: Tatra
- Argynnis euphrosyne f. answina Fruhstorfer, 1907; Ent. Zs. 21 (27): 164
- Argynnis euphrosyne fingal f. niveola Stichel, 1908; Berl. ent. Z. 53 (1): 81, pl. 3, f. 4
- Argynnis euphrosyne obscurior Seitz, 1909; Gross-Schmett. Erde 1: 229, pl. 67 h (Stgr. i.l.)
- Argynnis euphrosyne var. phaenna Jachontov, 1909; Revue Russe Ent. 8: 285; TL: Transcaucasia
- Argynnis euphrosyne cynosoma Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 15; TL: Salève
- Argynnis euphrosyne calynde Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 16; TL: Tessin, Fusio, Val Piora, 1250m
- Argynnis euphrosyne neston Fruhstorfer, 1917; Archiv Naturg. 82 A (2): 16; TL: Tessin, Mt Generoso
- interligata (Cabeau, 1919) (Argynnis); Rev. mens. Soc. ent. Namur. 1919: 49
- chlorographa (Cabeau, 1920) (Argynnis); Rev. mens. Soc. ent. Namur. 1920: 21; TL: France
- capronnieri (Derenne, 1922) (Argynnis); Rev. Mens. Soc. ent. Namur. 1922: 36
- rabesina (Cabeau, 1922) (Argynnis); Rev. mens. Soc. ent. Namur. 1922: 1
- Brenthis euphrosyne ab. plumbea Cockayne, 1924; Ent. Rec. J. Var. 36: 8; TL: Chilterns
- Brenthis euphrosyne alpina Ebert, 1926; Dt. ent. Z. Iris 40: 33
- = Clossiana euphrosyne lavinia ; [NHM card]
- Argynnis euphrosyne ab. pittionii Nitsche, 1926; Verh. zool.-bot. Ges. 74/75 (s.b.): 17
- forstorpensis (Strand, 1927) (Argynnis); Ent. Tidskr. 48: 56
- Boloria euphrosyne r. nestonclara Verity, 1932; Ent. Rec. J. Var. 44: 112; TL: Germany
- Boloria euphrosyne r. padimira Verity, 1932; Ent. Rec. J. Var. 44: 113; TL: Apennines
- Boloria euphrosyne r. eminens Verity, 1932; Ent. Rec. J. Var. 44: 113; TL: Ural Mts
- Boloria euphrosyne r. variana Verity, 1932; Ent. Rec. J. Var. 44: 114; TL: England
- Boloria euphrosyne r. magnaclara Verity, 1932; Ent. Rec. J. Var. 44: 120; TL: Amasia
- Boloria euphrosyne r. orientisvivax Verity, 1932; Ent. Rec. J. Var. 44: 121; TL: Sajan Mts
- vittata (Nordström, 1933) (Brenthis); Ent. Tidskr. 54: 175; TL: Lapland
- lecharlesi (Varin, 1933) (Argynnis); Amat. Papill. 6: 233
- septentrionalis (Nordström, 1933) (Brenthis); Ent. Tidskr. 54: 171; TL: Lapland
- Argynnis euphrosyne lavinia Hemming, 1934; Stylops 3 (9): 193 (repl. Brenthis euphrosyne alpina Ebert, 1926)
- dorensis (Réal, 1950) (Argynnis); Rev. franc. Lép. 12: 214; TL: France
- Clossiana euphrosyne r. austreminens Verity, 1950; Le Farfalle diurn. d'Italia 4: 240
- Clossiana euphrosyne neston sr. austroneston Verity, 1950; Le Farfalle diurn. d'Italia 4: 240
- = Clossiana euphrosyne neston ; [NHM card]
- Argynnis euphrosyne ab. edna Lobb, 1953; Ent. Rec. J. Var. 65: 56, pl. 3, f. 4-5; TL: Parkhurst Forest, Isle of Wight
- Brenthis euphrosyne esperi Heydemann, 1955; Scr. Naturw. Ver. Schleswig-Holstein 27 (2): 136 (Esper, 1777 1 (1) pl. 18)
- galiseri (Niculescu, 1957) (Argynnis); Comun. Acad. Repub. Rom. 7: 456
- albomaculata (Goodson, 1959) (Clossiana); Entomologist 92: 148
- Clossiana euphrosyne apennina; [NHM card]
- Clossiana euphrosyne answina; [NHM card]
- Clossiana euphrosyne austreminens; [NHM card]
- Clossiana euphrosyne calynde; [NHM card]
- Clossiana euphrosyne cynosoma; [NHM card]
- Clossiana euphrosyne eminens; [NHM card]
- Clossiana euphrosyne lavinia; [NHM card]
- Clossiana euphrosyne magnaclara; [NHM card]
- Clossiana euphrosyne nestonclara; [NHM card]
- Clossiana euphrosyne niveola; [NHM card]
- Clossiana euphrosyne neston; [NHM card]
- Clossiana euphrosyne orientisvivax; [NHM card]
- Clossiana euphrosyne obscurior; [NHM card]
- Clossiana euphrosyne pittionii; [NHM card]
- Clossiana euphrosyne padimira; [NHM card]
- Clossiana euphrosyne phaenna; [NHM card]
- Clossiana euphrosyne septentrionalis; [NHM card]
- Clossiana euphrosyne variana; [NHM card]
- Clossiana euphrosyne; [BOW]: pl. 1, f. 26; [EBW]; [BARU, #190]; [NHM card]; [BRU2]: 44, pl. 31, f. 13-24; [RFEL]
- Boloria euphrosyne; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
![[euphrosyne-1u.jpg]](_euphrosyne-1u.jpg)
500x349 (~71Kb) upperside Luckett, Cornwall, UK 20/05/95, Photo © Simon Coombes
|
![[euphrosyne-1v.jpg]](_euphrosyne-1v.jpg)
417x500 (~57Kb) underside Luckett, Cornwall, UK 20/05/95, Photo © Simon Coombes
|
![[euphrosyne-2l.jpg]](_euphrosyne-2l.jpg)
533x400 (~50Kb) larva An open pine grove above the bluff of the valley of the Bol'shoi Elbash River, 5 km NW of the village Ust'-Chem, Iskitim dist., Novosibirsk region, West Siberia, 9th May 1995 (mature larva), Photo © Oleg Kosterin
|
![[euphrosyne-3v.jpg]](_euphrosyne-3v.jpg)
600x832 (~62Kb) underside Finland, Loppi, Luutasuo 673:35, 2.6.1979, Photo © Pertti Pakkanen
|
![[euphrosyne-4fu.jpg]](_euphrosyne-4fu.jpg)
900x745 (~146Kb) upperside female FINLAND: Li: Inari, Luttojoki, 760:54, 11.7.1998, Photo © Markku Savela
|
![[euphrosyne-5u.jpg]](_euphrosyne-5u.jpg)
600x722 (~61Kb) upperside FINLAND, Ab: Suomusjärvi, 669:31, 15.6.1997, Photo © Markku Savela
|
![[euphrosyne-6.jpg]](_euphrosyne-6.jpg)
900x1291 (~125Kb) Finland, Ab: Suomusjärvi (669:31), 6.6.2004, Photo © Markku Savela
|
![[euphrosyne-7mu.jpg]](_euphrosyne-7mu.jpg)
1258x972 (~356Kb) upperside male An alpine meadow patch neighbouring with the most elevated larch wood outposts in the Kamdyt valley, a Tekelyu River right tributary which in turn is an Akkem River right tributary, the Katunskii Range northern principal slope, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Republic, West Siberia, Russia. 30th June 2007, Photo © Oleg Kosterin
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![[euphrosyne-7mv.jpg]](_euphrosyne-7mv.jpg)
861x1128 (~219Kb) underside male An alpine meadow patch neighbouring with the most elevated larch wood outposts in the Kamdyt valley, a Tekelyu River right tributary which in turn is an Akkem River right tributary, the Katunskii Range northern principal slope, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Rapublic, West Siberia, Russia. 2nd July 2007, Photo © Oleg Kosterin
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![[euphrosyne-8u.jpg]](_euphrosyne-8u.jpg)
779x555 (~89Kb) upperside Russia, Moscow area, 11.6.2011 (38°52'E, 55°33'N), Photo © D. Smirnov
|
![[euphrosyne-8v.jpg]](_euphrosyne-8v.jpg)
632x611 (~109Kb) underside Russia, Moscow area, 11.6.2011 (38°52'E, 55°33'N), Photo © D. Smirnov
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![[euphrosyne-v.jpg]](_euphrosyne-v.jpg)
800x631 (~57Kb) underside FINLAND: Ka: Virolahti, 671:53 (Reskopinsuo), 14.6.1997, Photo © Markku Savela
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Larva on
Viola ,
V. palustris,
V. riviniana,
Vaccinium uliginosum,
Ledum palustre [SPRK],
Viola hirta,
Waldsteinia ternata [BRU2], 45
-
B. e. euphrosyne CEU, Siberia
-
B. e. fingal (Herbst, 1800) NEU, Siberia
- euphrosyne fingal (Herbst, 1800) (Papilio); Naturs. Schmett. 10: pl. 270, f. 1-3
- Papilio lapponica Esper, 1798; Die Schmett., Suppl. Th 1 ([7]): 83, pl. 108, f. 5
- = Clossiana euphrosyne fingal ; [NHM card]
-
?B. e. nephele (Herrich-Schäffer, [1847]) Ural, Siberia
-
B. e. dagestanica (Sovinsky, 1905) Transcaucasia
- Clossiana euphrosyne dagestanica; [NHM card]
-
B. e. anka (Fruhstorfer, 1907) Caucasus
- Argynnis euphrosyne anka Fruhstorfer, 1907; Ent. Zs. 21 (27): 164; TL: Caucasus
- Clossiana phaenea Jachontov, 1908
-
B. e. orphana (Fruhstorfer, 1907) Transbaikalia, Amur, Ussuri
- Clossiana euphrosyne orphanus; [BARU]; [NHM card]
- Clossiana euphrosyne orphana; [BRU2], 45
-
B. e. rusalka (Fruhstorfer, 1909) SEU, W.Siberia
-
B. e. densoi (Fruhstorfer, 1909)
- Clossiana euphrosyne densoi; [NHM card]
-
B. e. kamtschadalis (Seitz, 1909) Kamchatka, N.Sakhalin
-
B. e. umbra (Seitz, 1909) Altai, Sayan
-
B. e. orphanoides (Huang & Murayama, 1992) Xinjiang
- Clossiana euphrosyne orphanoides Huang & Murayama, 1992; Tyô to Ga 43 (1): 7, f. 17; TL: Jiadengyu, Altai, 1380-1600m
-
Boloria iphigenia (Graeser, 1888)
-. See [maps]
- Argynnis iphigenia Graeser, 1888; Berl. Ent. Zs. 32 (1): 90; TL: Amur "downstream of Khabarovsk"
Larva on
Viola selkirkii,
(mainland,Japan) Viola grypoceras,
(Sakhalin,Japan) [BARU] [BRU2], 45
-
B. i. iphigenia E.Amur, Ussuri, NE.China
-
B. i. sachalinensis (Matsumura, 1908) Japan
- Argynnis euphrosyne var. sachalinensis; Matsumura, 1919, Thous. Ins. Japan. Addit. 3: 585, pl. 45, f. 10
- Argynnis euphrosyne sachalinensis; Esaki, 1916, Ent. Mag. Kyoto 2 (1): 48, 50, pl. 2, f. 6; Matsumura, 1925, J. Coll. Agric. Hokkaido imp. Univ. 15: 99
- Argynnis selene sachalinensis; Nire, 1916, Zool. Mag. Tokyo 28: 64
- Clossiana euphrosyne sachalinensis; [NHM card]
- Boloria selene sachalinensis; [NHM card]
- Clossiana iphigenia sachalinensis; Kudrna, 1974, Atalanta 5: 108; [NHM card]
- ?Clossiana iphigenia sachalinensis; [BARU]
-
B. i. insularia (Korshunov, 1996) Sakhalin, Kunashir
- Clossiana iphigenia alpharatoria; [BRU2], 45
- Clossiana iphigenia insularia; [RFEL]
- Boloria iphigenia insularia; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 32
-
Boloria kriemhild (Strecker, 1879)
Montana, Wyoming. See [maps]
Relict Fritillary
Kriemhild Fritillary
- Argynnis kriemhild Strecker, 1879; in McCauley, Rept. San Juan Recon. 5 (1) pt. 2 (3) appendix SS: 1854, pl. 1, f. 5-6
- Argynnis laurenti Skinner, 1913; Ent. News 24 (10): 450; TL: Silver Lake, Utah
- Clossiana lücki [= luecki] Reuss, 1923; Soc. Ent. 38 (4): 14; TL: Utah
- Clossiana kriemeld; McHenry, 1964, (missp.)
- Argynnis laurenti; [NHM card]
- Clossiana lucki; dos Passos, 1964, (missp.)
- Clossiana lucki[sic, recte luecki]; [NHM card]
- Brenthis epithore kreimhild[sic]; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
- Clossiana kriemhild; [NACL], #4468
- Boloria epithore kriemhild; [NHM card]
- Clossiana euphrosyne kriemhild; [NHM card]
- Boloria kriemhild; [BNA]: 320, #241, pl. 26; [Opler]
- Boloria (Clossiana) kriemhild; Pelham, 2008, J. Res. Lepid. 40: 301
Larva on
Viola [NAB]
-
Boloria freija (Thunberg, 1791)
. See [maps]
Muurainhopeatäplä
Freija's Fritillary
Freya's Fritillary
Zigzag Fritillary
Frejas pärlemorfjäril
Myrpärlefjäril
- ?Clossiana freija daisetuzana; [NHM card]
- Brenthis freija; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
- Clossiana freija; [BARU, #201]; [NACL], #4471; [MRS], 477; [NHM card]; [BRU2]: 48, pl. 32, f. 24-29; [RFEL]
- Boloria freija; [BNA]: 323, #249, pl. 26; [BOC], 284; [Opler]; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
- Boloria (Clossiana) freija; Pelham, 2008, J. Res. Lepid. 40: 304
Larva on
Rubus chamaemorus,
Vaccinium uliginosum [SPRK],
Vaccinium vitis-idaeus,
Rhododendron auerum [BARU],
Empetrum nigrum [MRS],
Vaccinium caespitosum,
V. uliginosum,
Arctostaphylos uva-ursi,
Empetrum nigrum [NAB],
Arctous alpina,
(Scandinavia) Sieversia ,
Rhododendron ,
(Japan) Dryas integrifolia,
(N.America) [BRU2], 48
-
B. f. freija Fennoscandia, W.Siberia, C.Siberia, Far East, Transbaikiali, Chukotka, Amur, Ussuri, Canada, Alaska
- Argynnis freya Godart, 1819; Encyclopédie Méthodique 9 (1): 273 (repl. Argynnis freija Thunberg, 1791)
- Papilio freija Quensel, 1791; K. VetenskAcad. Handl. [2] 12: 276, pl. 10, f. 5-6 (preocc. Papilio freija Thunberg, 1791)
- Papilio lapponica Esper, 1789; Die Schmett., Suppl. Th 1 (3-4): 21, pl. 97, f. 3
- Argynnis (Bretis[sic]) freija jakutensis Wnukowsky, 1927; Mitt. Münch. Ent. Ges. 17 (7-12): 70; TL: Jakutia
- Brenthis freija tr.f. gunderi Harper, 1933; Pan-Pacific Ent. 9 (3): 99; TL: McCreary, Manitoba, Canada
- Clossiana freija zamolodchikova Churkin, 2001; Helios 2: 99, pl. 8, f. 10-12
-
B. f. tarquinius (Curtis, 1835) British Columbia, ...
-
B. f. pallida (Elwes, 1899) Altai, Sayan, Tuva
![[pallida-1fv.jpg]](_pallida-1fv.jpg)
533x400 (~67Kb) underside female A dwarf birch (Betula rotundifolia Spach) montane tundra at 2200 m above sea level in the Argem (Direntai) Stream basin, 15 km W of the village Dzhazatr, on the eastern spurs of the Katunskiy Mt. Range, Kosh-Agach District, Central Altai Mts., West Siberia, Russia. 13th July, 1988, Photo © Oleg Kosterin
|
![[pallida-1mu.jpg]](_pallida-1mu.jpg)
508x444 (~38Kb) upperside male A meadowy-tundrous SW slope of a ridge between the headwaters of the Chikty Rivulet, 2600 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 10th July 1998 (on Aster alpinus), Photo © Oleg Kosterin
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![[pallida-1xmu.jpg]](_pallida-1xmu.jpg)
600x600 (~71Kb) upperside male A meadowy-tundrous SW slope of a ridge between the headwaters of the Chikty Rivulet, 2600 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 10th July 1998 (on Aster alpinus), Photo © Oleg Kosterin
|
-
B. f. asahidakeana (Matsumura, 1926) Japan
- Argynnis asahidakeana; [NHM card]
- Clossiana freija asahidakeana; Kudrna, 1974, Atalanta 5: 109
-
B. f. browni (Higgins, 1953) Rocky Mountains
- Clossiana freija browni Higgins, 1953; Entomologist 86 (9): 210
-
Boloria natazhati (Gibson, 1920)
Yukon, Victoria I.. See [maps]
Pleistocene Fritillary
- Boloria freija natazhati; dos Passos, 1964; Howe, 1975
- Clossiana freija natazhati; Miller & Brown, 1981; Ferris, dos Passos, Ebner & Lafontaine, 1983; [NACL], #4471b; Tilden & Smith, 1968
- Clossiana natazhati; [NHM card]
- Boloria natazhati; [BNA]: 323, #250, pl. 24; Troubridge & Wood, 1990, J. Lep. Soc. 44 (3): 181, f. 1-2; [Opler]
- Boloria (Clossiana) natazhati; Pelham, 2008, J. Res. Lepid. 40: 304
Larva on
Dryas integrifolia? Troubridge & Wood, 1990, J. Lep. Soc. 44 (3): 185
-
-
B. n. nabokovi Stallings & Turner, 1947 Alaska
- Boloria freija nabokovi Stallings & Turner, 1947; Can. Ent. 78 (7-8): 134; TL: Alaska Military Highway, 102 miles N of Summit 2, 1830m
- Boloria freija nabokovi; dos Passos, 1964; Howe, 1975
- Clossiana freija nabokovi; Miller & Brown, 1981; [NACL], #4471c; Tilden & Smith, 1986; [NHM card]
- Boloria natazhati nabokovi; Pelham, 2008, J. Res. Lepid. 40: 305
-
B. n. bankslandia Shepard, 1998 Victoria Is.
-
Boloria angarensis (Erschoff, 1870)
-. See [maps]
![[angarensis-1fu.jpg]](_angarensis-1fu.jpg)
533x400 (~45Kb) upperside female An open larch (Larix gmelinii (Rupr.) Rupr.)/birch (Betula pendula Roth. and B. davurica Pallas) forest above the left bluff of the Budyumkan River valley 5 km of its confluence with the Argun' River 6 km S of the village Uryupino, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 2nd August 1997, Photo © Oleg Kosterin
|
![[angarensis-1fv.jpg]](_angarensis-1fv.jpg)
352x299 (~22Kb) underside female A road in a larch (Larix gmelinii (Rupr.) Rupr.) forest not far from the Argun' River left bank between the village Uryupino and the Gazimur River, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 31th July 1997, Photo © Oleg Kosterin
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Larva on
Vaccinium [BRU2], 43
Larva on
Viola [RFEL]
-
B. a. angarensis Transbaikalia, S.Siberia
-
B. a. herzi (Wnukowsky, 1927) N.Transbaikalia, Far East Yakutia
-
B. a. samkoi (Sheljuzhko, 1931) Polar Urals, Yamal Peninsula
-
B. a. sedychi (Weiss, 1964) Polar Urals, Yamal Peninsula
-
B. a. alticola (Sushkin & Tshetverikov, 1907) Sayan, Tuva (mountains)
-
B. a. miyakei (Matsumura, 1919) Sakhalin
-
B. a. hakutozana (Matsumura, 1927) Amur, Ussuri, N.Korea, NE.China
- Argynnis hakutozana; [NHM card]
- Clossiana hakutozana; [BARU, #195]
- Clossiana hakutosana[sic, recte hakutozana]; [EBW], (name only)
- Clossiana angarensis hakutozana; [BRU2], 43; [RFEL]
- Boloria angarensis hakutozana; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
-
Boloria thore (Hübner, [1803-1804])
Japan. See [maps]
Purohopeatäplä
Thore's Fritillary
Gråkantad pärlemorfjäril
Bäckpärlefjäril
Alpen-Perlmutterfalter
![[thore-1u.jpg]](_thore-1u.jpg)
600x600 (~98Kb) upperside forest-tundra belt of the Koni Peninsula, S Magadan Region, Russia. 11th of July 1989, Photo © Oleg Kosterin
|
![[thore-1v.jpg]](_thore-1v.jpg)
374x506 (~29Kb) underside Among the dead branches of the Siberian dwarf pine (Pinus pumila Ledeb.) A forest/tundra on the Koni Peninsula, S Magadan Region, Russia. 11th of July 1989, Photo © Oleg Kosterin
|
![[thore-u.jpg]](_thore-u.jpg)
494x684 (~70Kb) upperside
|
![[thore-v.jpg]](_thore-v.jpg)
470x733 (~69Kb) underside
|
|
Larva on
Viola bifolia,
(Finland) Viola altaica,
(Urals) [BRU2], 47 Viola selkirkii,
Viola grypoceras [MRS]
-
B. t. thore Alps
- Clossiana thore thore; [BRU2], 46
-
B. t. borealis (Staudinger, 1861) N.Scandinavia, N.Siberia, Far East, Kamchatka
-
B. t. transuralensis (Sheljuzhko, 1931) Ural, W.Siberia
- Brenthis thore transuralensis Sheljuzhko, 1931; Int. ent. Zs. 25: 46; TL: "Umgegend von Tobolsk"
-
B. t. hyperlampra (Fruhstorfer, 1907) Pyrenees
- Clossiana thore hyperlampra; [NHM card]
-
B. t. hypercala (Fruhstorfer, 1907) Altai, Sayan, Transbaikalia
![[hypercala-1fu.jpg]](_hypercala-1fu.jpg)
600x600 (~102Kb) upperside female An open spruce forest, the Stolby Nature Reserve, Krasnoyarsk Province, 11th July 1995, Photo © Oleg Kosterin
|
![[hypercala-2fum.jpg]](_hypercala-2fum.jpg)
784x1098 (~137Kb) upperside male female A grassy open larch forest on the Yabaganskii Pass (1400 m above sea level) through the Seminskii mountain range, the northern Altai Mts., West Siberia, Russia. 11th July 1999, Photo © Oleg Kosterin
|
![[hypercala-2fv.jpg]](_hypercala-2fv.jpg)
800x800 (~145Kb) underside female A grassy open larch forest on the Yabaganskii Pass (1400 m above sea level) through the Seminskii mountain range, the northern Altai Mts., West Siberia, Russia. 11th July 1999, Photo © Oleg Kosterin
|
![[hypercala-2mv.jpg]](_hypercala-2mv.jpg)
1088x786 (~121Kb) underside male A grassy open larch forest on the Yabaganskii Pass (1400 m above sea level) through the Seminskii mountain range, the northern Altai Mts., West Siberia, Russia. 11th July 1999, Photo © Oleg Kosterin
|
|
-
B. t. herculea Fruhstorfer, 1907² ?
- thore herculea Fruhstorfer, 1907²
- herculea (Fruhstorfer, 1907); ; TL: Altai
- Clossiana thore herculea; [BARU]
-
B. t. hyperusia (Fruhstorfer, 1907) Amur, Ussuri
- Argynnis thore hyperusia Fruhstorfer, 1907; Ent. Zs. 21 (27): 164; TL: Amur
- = Clossiana thore borealis ; [RFEL]
- Argynnis thore meridionalis Kardakoff, 1928; Ent. Mitt. 17 (4): 270, pl. 7, f. 5-6
- = Clossiana thore brenda ; [NHM card]
- = Clossiana thore hyperusia ; [BRU2], 47
- Clossiana brenda Hemming, 1933; Entomologist 66 (12): 276 (repl. Argynnis thore meridionalis Kardakoff, 1928)
- = Clossiana thore hyperusia ; [BRU2], 47
-
B. t. excellens (Krulikovsky, 1893)
- Argynnis thore var. excellens Krulikovsky, 1893; Soc. Ent. 7 (21) (24): 187; TL: Rossia orientalis
- splendida (Krulikovsky, 1895)
-
B. t. karafutonis (Matsumura, 1919) Sakhalin
-
Boloria frigga (Thunberg, 1791)
. See [maps]
Rahkahopeatäplä
Frigga's Fritillary
Willow-Bog Fritillary
Friggas pärlemorfjäril
Hjortronpärlefjäril
- Papilio frigga Thunberg, 1791; Diss. Ent. sistens Insecta Suecica (2): 33 (& Becklin ?); TL: Lappland
- Argynnis frigga ab. ahti Sahlberg, 1891; Medd. Soc. Fenn. 18: 205, f. 4; TL: Härkäsuo, Sammatti
- Argynnis frigga ab. tapio Sahlberg, 1891; Medd. Soc. Fenn. 18: 205, f. 5; TL: Härkäsuo, Sammatti
- Argynnis frigga f. ahti Stichel, 1908; Berl. ent. Z. 53 (1): 86, pl. 3, f. 7
- Argynnis frigga f. tapio Stichel, 1908; Berl. ent. Z. 53 (1): 86, pl. 3, f. 8
- Brenthis frigga lapponica Lingonblad, 1944; Notulae Ent. 24 (1-2) (1-2): 51; TL: Finland, Lappland
- Brenthis (Clossiana) frigga australis Lingonblad, 1946; Notulae Ent. 26 (1-2): 55 (preocc. Argynnis oscarus var. australis Graese, 1888); TL: Finland, Muonio, Enontekiö
- exmaculata (Roos, 1953) (Clossiana); Opusc. ent. 18: 229
- Clossiana frigga ahti; [NHM card]
- Clossiana frigga australis; [NHM card]
- Clossiana frigga tapio; [NHM card]
- Brenthis frigga; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
- Clossiana frigga; [NACL], #4466; [BARU, #196]; [NHM card]; [BOW]: pl. 1, f. 27; [BRU2]: 47, pl. 32, f. 13-23; [RFEL]
- Boloria frigga; [BNA]: 321, #244, pl. 26; [Opler]; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
- Boloria (Clossiana) frigga; Pelham, 2008, J. Res. Lepid. 40: 300
![[frigga-u.jpg]](_frigga-u.jpg)
461x381 (~37Kb) upperside
|
![[frigga-v.jpg]](_frigga-v.jpg)
447x373 (~36Kb) underside
|
Larva on
Rubus chamaemorus [SPRK],
Salix ,
Betula ,
Dryas integrifolia [BNA], 321
-
B. f. frigga N.Scandinavia
- Boloria frigga helvenacius Sedykh, 1977; Geogr. Aspect. Conserv. Flora Faun. NE Europ. USSR: 102; TL: Polar Urals
-
B. f. saga (Staudinger, 1861) Canada
- Argynnis frigga var. saga Staudinger, 1861; Stettin Ent. Ztg 22 (10-12): 350 (Kaden); TL: Labrador
- Brenthis frigga saga Kaden, 1861; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
- Argynnis friggar var. saga Kaden, 1861; Strecker, 1878, Synon. Cat. Macroplep.: 117
- Clossiana frigga saga; [NACL], #4466a; [NHM card]
- Boloria frigga saga; [BNA], #244; Pelham, 2008, J. Res. Lepid. 40: 300
-
B. f. alpestris (Elwes, 1899) Altai, Sayan, Transbaikalia
![[alpestris-1fu.jpg]](_alpestris-1fu.jpg)
872x600 (~56Kb) upperside female Highland dwarf birch tundras on of the southerm principal slope of the Yuzhno-Chuiskii mountain at the the Chikty Rivulet headwaters, 2400 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 15th July 1998, Photo © Oleg Kosterin
|
![[alpestris-1fux.jpg]](_alpestris-1fux.jpg)
900x592 (~98Kb) upperside female A flowery alpine meadow in a valley of the rivulet descending from the southerm principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets, 2600 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 11th July 1998, Photo © Oleg Kosterin
|
![[alpestris-1fvx.jpg]](_alpestris-1fvx.jpg)
800x800 (~121Kb) underside female A flowery alpine meadow in a valley of the rivulet descending from the southerm principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets, 2600 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 11th July 1998, Photo © Oleg Kosterin
|
![[alpestris-1mv.jpg]](_alpestris-1mv.jpg)
475x500 (~33Kb) underside male An alpine meadow vegetation at a brook flowing in a moss/sedge tundra in a cirque of the sources of a rivulet descending from the southerm principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets, 2650 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 16 July 1998, Photo © Oleg Kosterin
|
|
-
B. f. annae (Suschkin, 1906) Tarbgatai Mts.
-
B. f. sagata (Barnes & Benjamin, 1923) Wyoming - Colorado
- Brenthis frigga r. sagata Barnes & Benjamin, 1923; Can. Ent. 55 (6): 146; TL: Hall Valley, Colorado
-
B. f. gibsoni (Barnes & Benjamin, 1926) Alaska - Hudson Bay
- Brenthis frigga r. gibsoni Barnes & Benjamin, 1926; Bull. South. Calif. Acad. Sci. 25 (3): 92; TL: Barter I., Northern Alaska
- Argynnis frigga f. alaskensis Lehmann, 1913; Gross-Schmett. Erde 5: 424, pl. 87 e (preocc. Brenthis pales var. alaskensis Holland, 1900); TL: NE.Alaska, 69°40N, 141°W
- Brenthis frigga var. lehmanni Holland, 1928; Ann. Carnegie Mus. 19 (1): 44 (repl. Argynnis frigga f. alaskensis Lehmann, 1913); TL: NE.Alaska, 69°40N, 141°W
- Boloria frigga kosarevi Korb, 2011; Eversmannia Suppl. 2: 33 (repl. Argynnis frigga f. alaskensis Lehman, 1913); TL: NE.Alaska, 69°40N, 141°W
-
B. f. maritima (Kardakoff, 1928) Amur, Ussuri
- Argynnis frigga maritima Kardakoff, 1928; Ent. Mitt. 17 (4): 270, pl. 7, f. 4; TL: Primorye, Ternei Bay
-
Boloria dia (Linnaeus, 1767)
-. See [maps]
Violet Fritillary
Hainveilchen-Scheckling
La Petite Violette
Perlada Violeta
Dostojka dia
- Papilio dia Linnaeus, 1767; Syst. Nat. (Edn 12) 1 (2): 785; TL: Austria
- Papilio dia minor Borkhausen, 1788; Naturges. Eur. Schmett. 1: 41
- Brenthis nenoquis Reakirt, 1866; Proc. Acad. nat. Sci. Philad. 18 (3): 247; TL: California [error]
- Argynnis dia ab. vittata Spuler, 1901; Schmett. Eur. 1: 28
- Argynnis dia ab. nigrostriata Schultz, 1908; Ent. Zs. 22 (9): 39
- Argynnis dia ab. senta Schultz, 1908; Ent. Zs. 22 (10): 40
- Argynnis dia ab. mediofasciata Schultz, 1908; Soc. Ent. 22 (23): 177
- Argynnis dia diniensis Oberthür, 1909; Étud. Lépid. Comp. 3: 216
- Argynnis dia ab. ornata Aigner-Abafi, 1906; Ann. hist.-nat. Mus. Hung. 4 (2): 503
- Argynnis dia ab. hudaki Aigner-Abafi, 1906; Rovart. Lapok 13 (4): 72
- Argynnis dia ab. hudaki Aigner-Abafi, 1906; Ent. Zs. 19 (35): 208; TL: Vinkovece
- Argynnis dia ab. lugens Grund, 1908; Soc. ent. 23 (11): 82
- nigricans (Ksienschopolski, 1911) (Argynnis); Trudy obshch.Izsl. Volyni Zitomir 8: 40, pl. 1, f. 5
- Brenthis dia r. laetior Verity, 1919; Ent. Rec. J. Var. 31: 194
- Brenthis dia f. flavens Verity, 1919; Ent. Rec. J. Var. 31: 194
- interligata (Cabeau, 1922) (Argynnis); Rev. mens. Soc. ent. Namur. 1922: 22; TL: France
- ?euphrosynella (Cabeau, 1924) (Argynnis); Rev. mens. Soc. ent. Namur. 24: 34
- bivittata (Cabeau, 1925) (Argynnis); Rev. mens. Soc. ent. Namur. 25: 7; TL: Belgium
- posteronubilata (Cabeau, 1930) (Argynnis); Lambillionea 30: 122
- Boloria dia f. postdia Verity, 1933; Bull. Soc. ent. Fr. 38 (6): 74
- Argynnis dia f. albomaculata Bedrnik, 1936; Polsk. Pismo ent. 13: 197
- couleti (Oberthür, 1937) (Argynnis); Lambillionea 37: 26; TL: Basses-Alpes
- reducta (Lumma, 1938) (Argynnis); Ent. Zs. 52 (13): 104
- Clossiana postdia; Verity, 1950, Le Farfalle diurn. d'Italia 4: 247
- Clossiana dia albomaculata; [NHM card]
- Argynnis dia diniensis; Fruhstorfer, 1917, Archiv Naturg. 82 A (2): 18
- Clossiana dia; [BOW]: pl. 1, f. 28; [EBW]; [BARU, #200]; [MRS], 477; [NHM card]; [BRU2]: 49, pl. 33, f. 13-21; [RFEL]
- Boloria dia; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
![[dia-1v.jpg]](_dia-1v.jpg)
533x400 (~44Kb) underside A dwarf birch (Betula rotundifolia Spach) montane tundra at 2200 m above sea level in the Argem (Direntai) Stream basin, 15 km W of the village Dzhazatr, on the eastern spurs of the Katunskiy Mt. Range, Kosh-Agach District, Central Altai Mts., West Siberia, Russia. 129 - 15th, 130 - 12th July, 1988, Photo © Oleg Kosterin
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![[dia-2l.jpg]](_dia-2l.jpg)
598x400 (~44Kb) larva Central Europe 1990-2000, Photo © Michael Borsch
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![[dia-2u.jpg]](_dia-2u.jpg)
598x400 (~45Kb) upperside Central Europe 1990-2000, Photo © Michael Borsch
|
![[dia-3u.jpg]](_dia-3u.jpg)
1100x768 (~335Kb) upperside Russia, West Siberia, Altaiskii Krai Province, Zmeinogorsk District, Savvushka village environs, the Lake Kolyvenskoe bank, a slope steppe. 9th September 2007, Photo © Oleg Kosterin
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![[dia-u.jpg]](_dia-u.jpg)
465x500 (~43Kb) upperside Axat, Pyrenees orientales, France 24/07/96, Photo © Simon Coombes
|
![[dia-v.jpg]](_dia-v.jpg)
500x503 (~50Kb) underside Axat, Pyrenees orientales, France 24/07/96, Photo © Simon Coombes
|
Larva on
Viola ,
Rubus idaeus,
Prunella vulgaris [BRU2], 49
-
-
B. d. nigrescens (Schavrov, 1886)
- dia nigrescens (Schavrov, 1886) (Argynnis); ? 50: 203
-
B. d. leonina (Fruhstorfer, 1909)
-
B. d. setania (Fruhstorfer, 1909)
- Clossiana dia setania; [BMAT]: 105, pl. 32, f. 19-20 (not in N.Africa); [NHM card]
-
B. d. disconota (Krulikovsky, 1909) CEU, W.Siberia
-
B. d. calida (Jachontov, 1911) Caucasus, Transcaucasus
- Argynnis dia var. calida Jachontov, 1911; Mitt. Kaukas. Mus. 5: 306
-
B. d. alpina (Elwes, 1899) Dzhungarsky Alatau, Altai, Sayan, Transbaikalia
-
B. d. semota (Tuzov, 1999) Magadan
- Clossiana dia semota Tuzov, 1999; Helios 1: 105; TL: Russia, Magadan
-
Boloria gong (Oberthür, 1884)
Tibet, W.China, N.China. See [maps]
-
B. g. charis (Oberthür, 1891) Yunnan
-
?B. g. splendens (Murayama, 1983)
- ?Clossiana gong splendens; [MRS], 477
-
B. g. xizangensis (Huang, 2000) SE.Tibet
- Clossiana gong xizangensis Huang, 2000
-
Boloria titania (Esper, 1794)
. See [maps]
Lehtohopeatäplä
Titania's Fritillary
Purple Bog Fritillary
Amathusias pärlemofjäril
Lundpärlefjäril
Natterwurz-Perlmutterfalter
Dostojka titania
- Papilio titania Esper, 1794; Die Schmett., Suppl. Th 1 (3-4): 58, (5-6) pl. 103, f. 4; TL: Sardinia
- Papilio dia major Esper, 1786; Die Schmett. Th. I, Bd. 2 (28): pl. 93, f. 2-3
- Papilio diana Hübner, [1799-1800]; Samml. eur. Schmett. [1]: pl. 10, f. 51-54
- Argynnis amathusia ab. radiifera Schultz, 1908; Soc. Ent. 22 (23): 177
- Argynnis amathusia ab. nigrata Schultz, 1908; Soc. Ent. 22 (23): 177
- Argynnis amathusia var. transsylvanica Tiltscher, 1913; Ent. Zs. 26: 210
- Argynnis amathusia ab. bialoviezensis Gieysztor, 1923; Bialowieza 2: 125
- Boloria amathusia titania f. microtitania Verity, 1933; Ent. Rec. J. Var. 45 (6): 91
- Argynnis amathusia nansetsuzana Doi, 1935; Zephyrus 6: 15; TL: Korea
- lemagneni (Plantrou, 1960) (Clossiana); Alexanor 1: 199
Misidentified/misapplied:
- Clossiana titania (= Clossiana chariclea); [NACL], #4474
- Boloria titania (= Clossiana chariclea); [BNA]: 319, #240
![[titania-1a.jpg]](_titania-1a.jpg)
571x653 (~100Kb) Russia, Moscow area (36°25'E, 56°00'N), 24.7.2009, Photo © D. Smirnov
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![[titania-1v.jpg]](_titania-1v.jpg)
604x640 (~97Kb) underside Russia, Moscow area (36°25'E, 56°00'N), 24.7.2009, Photo © D. Smirnov
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Larva on
Polygonum ,
Filipendula ,
Trollius ,
Viola [RFEL]
-
B. t. titania Sardinia, ?
-
B. t. cypris (Meigen, 1829) S.Germany, Switzerland
-
B. t. bivina (Fruhstorfer, 1908) CEU, SEU
Larva on
Viola ,
Vaccinium uliginosum [BRU2], 50
-
B. t. bosna (Fruhstorfer, 1908) Bosnia
- Clossiana titania bosna; [NHM card]
-
B. t. dinara (Fruhstorfer, 1908) Herzegowina
- Clossiana titania dinara; [NHM card]
-
B. t. serena (Fruhstorfer, 1908) S.Tirol
- Argynnis amathusia serena Fruhstorfer, 1908; Int. ent. Zs. 2 (33): 214; TL: Wallis, Zermatt and Simplon region, S.Tirol, Piemont, Courmayeur
- Clossiana titania serena; [NHM card]
-
B. t. pralognana (Fruhstorfer, 1908)
- Clossiana titania pralognana; [NHM card]
-
B. t. halesa (Fruhstorfer, 1908) Pyrenees
- Clossiana titania halesa; [NHM card]
-
B. t. blachieri (Fruhstorfer, 1909)
- Clossiana titania blachieri; [NHM card]
-
B. t. jugurtha (Fruhtorfer, 1909)
- Argynnis amathusia jugurtha Fruhstorfer, 1909; Int. ent. Zs. 3 (4): 20; TL: Baeyern, Kochelsee, Berchesgaed, Triol, Karrersee
- Clossiana titania jugurtha; [NHM card]
-
B. t. staudingeri (Wnukowsky, 1929) S.Siberia, Altai, Sayan, mountains (Tuva), Transbaikalia, Amur, Sakhalin?
- titania staudingeri (Wnukowsky, 1929) (Brenthis); Zool. Anz. 83 (9/10): 222 (repl. Argynnis amathusia var. sibirica Staudinger, 1892); TL: Kentei
- Argynnis amathusia var. sibirica Staudinger, 1892; Dt. Ent. Z. Iris 5 (2): 330 (preocc. Argynnis selenis var. sibirica Erschoff, 1870); TL: Kentei
- = Clossiana titania staudingeri ; [NHM card]
- Argynnis amathusia f. altaica Seitz, 1909; Gross-Schmett. Erde 1: 232, pl. 68 f (preocc. Argynnis pales var. altaica Grum-Grshimailo, 1893); TL: Kentei
- = Clossiana titania julia ; [NHM card]
- Argynnis titania tana Hemming, 1934; Stylops 3 (9): 193 (repl. Argynnis amathusia f. altaica Seitz, 1909)
- = Clossiana titania julia ; [NHM card]
- Argynnis titania julia Hemming, 1934; Stylops 3 (5): 97 (repl. Argynnis amathusia f. altaica Seitz, 1909)
![[staudingeri-1u.jpg]](_staudingeri-1u.jpg)
700x510 (~58Kb) upperside A grassy open larch forest on the Yabaganskii Pass (1400 m above sea level) through the Seminskii mountain range, the northern Altai Mts., West Siberia, Russia. 11th July 1999, Photo © Oleg Kosterin
|
Larva on
Bistorta major,
Filipendula ulmaria,
Trollius asiaticus [BRU2], 50
-
B. t. minuscula (Eisner, 1942) Lithuania
- Clossiana titania minuscula; [NHM card]
-
Boloria chariclea (Schneider, 1794)
. See [maps]
Tundrahopeatäplä
Arctic Fritillary
Tundrapärlefjäril
- Papilio chariclea Schneider, 1794; Neuest. Mag. Lieb. Ent. 1 (5): 588; TL: Lappland
- aili (Kontuniemi, 1939) (Argynnis); Ann. ent. fenn. 5: 165
- Clossiana carichlea; Zetterstedt, 1839, (missp.)
- Clossiana churiclea; Johansen, 1921, (missp.)
- Clossiana churchilea; dos Passos, 1964, (missp.)
- Brenthis chariclea; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
- Clossiana chariclea; [NACL], #4475; [BARU, #199]; [BOW]: pl. 1, f. 25; [NHM card]; [BRU2]: 48, pl. 33, f. 4-9; [RFEL]
- Boloria chariclea; [BOC], 286; [Opler]; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
- Boloria (Clossiana) chariclea; Pelham, 2008, J. Res. Lepid. 40: 305
![[chariclea-1u.jpg]](_chariclea-1u.jpg)
550x424 (~48Kb) upperside FINLAND: Li: Utsjoki Puollamoaivi, 775:50, 8.7.1997, Photo © Markku Savela
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![[chariclea-1v.jpg]](_chariclea-1v.jpg)
500x564 (~35Kb) underside FINLAND: Li: Utsjoki Ailigas, 775:50, 8.7.1997, Photo © Markku Savela
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![[chariclea-2u.jpg]](_chariclea-2u.jpg)
700x506 (~69Kb) upperside FINLAND: Li: Utsjoki Ailigas, 775:50, 6.7.1997, Photo © Markku Savela
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Larva on
Dryas integrifolia,
Salix arctica,
S. reticulata,
(N.America) [BRU2], 49
-
Boloria bellona (Fabricius, 1775)
Rocky Mountains, Canada, Washington, Oregon. See [maps]
Meadow Fritillary
Misidentified/misapplied:
Larva on
Viola sororia,
V. pallens [BNA], 320
-
-
B. b. toddi (Holland, 1928) Labrador, N.Manitoba, Northwest Territories
-
B. b. jenistai Stallings & Turner, 1947
- Boloria toddi jenistai Stallings & Turner, 1947; Can. Ent. 78 (7-8): 135; TL: Lloydminster, Sask.
-
Boloria epithore (Edwards, 1864)
Yukon - California. See [maps]
Western Meadow Fritillary
Pacific Fritillary
![[epithore-1uU.jpg]](_epithore-1uU.jpg)
identification uncertain 600x539 (~44Kb) upperside USA: Near Crystal Mtn, (46°57'10N 121°28'27W), Pierce Co., WA, 27.7.1999, Photo © Markku Savela
|
![[epithore-1vU.jpg]](_epithore-1vU.jpg)
identification uncertain 400x369 (~20Kb) underside USA: Near Crystal Mtn, (46°57'10N 121°28'27W), Pierce Co., WA, 27.7.1999, Photo © Markku Savela
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![[epithore-2fu.jpg]](_epithore-2fu.jpg)
1000x860 (~111Kb) upperside female USA: Washington, Kittitas Co., Tucquala Lake, 24.7.2011, Photo © Markku Savela
|
![[epithore-2v.jpg]](_epithore-2v.jpg)
1000x837 (~96Kb) underside USA: Washington, Kittitas Co., Tucquala Lake, 24.7.2011, Photo © Markku Savela
|
![[epithore-3u.jpg]](_epithore-3u.jpg)
1400x1046 (~252Kb) upperside USA: Washington, Skagit Co., on Sauk Mountain Rd (48°31'23N 121°36'22W), 15.8.2012, Photo © Markku Savela
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![[epithore-3v.jpg]](_epithore-3v.jpg)
1000x925 (~153Kb) underside USA: Washington, Skagit Co., on Sauk Mountain Rd (48°31'23N 121°36'22W), 15.8.2012, Photo © Markku Savela
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![[epithore-4a.jpg]](_epithore-4a.jpg)
1000x1249 (~153Kb) USA: Washington, Skagit Co., Sauk Mountain Trail Head & Trail, 15.8.2012, Photo © Markku Savela
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![[epithore-4u.jpg]](_epithore-4u.jpg)
1200x957 (~152Kb) upperside USA: Washington, Skagit Co., Sauk Mountain Trail Head & Trail, 15.8.2012, Photo © Markku Savela
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|
Larva on
Viola sempervirens,
V. glabella,
V. ocellata,
V. adunca,
V. nephrophylla? [BOC], 283
-
B. e. epithore California
-
B. e. chermocki Perkins & Perkins, 1966 Oregon
- Boloria epithore chermocki Perkins & Perkins, 1966; J. Lep. Soc. 20 (2): 109; TL: 2.9 mi E of Dolph, Yamhill Co., Oregon
- Boloria epithore uslui Koçak, 1984; Priamus 3 (3): 96 (repl. Boloria epithore borealis Perkins, 1973); TL: Shingle Creek Rd., Keremeos, BC
- Boloria epithore borealis Perkins, 1973; Bull. Allyn Mus. 11: 6, pl. 1, f. 13-16 (preocc. Argynnis thore borealis Staudinger, 1861); TL: Shingle Creek Rd., Keremeos, BC
- Brenthis epithore ab. ♀ obscuripennis Gunder, 1926; Ent. News 37 (1): 7; TL: Chilcolin, Br. Columbia
-
B. e. sierra Perkins, 1973 California
- Boloria epithore sierra Perkins, 1973; Bull. Allyn Mus. 11: 10, pl. 1, f. 1-3; TL: Sentinel Dome, Yosemite Nat. Park, California
- Brenthis epithore ab. eldorado Strand, 1915; Arch. Naturgesch. 80 A (11): 156; TL: Plumas Co. ... Eldorado Co., California
- Brenthis epithore ab. wawonae Gunder, 1924; Ent. News 35 (5): 156; TL: Wawona, Mariposa Co., California
-
B. e. sigridae (Shepard, 2001) British Columbia
- Clossiana epithore sigridae Shepard, 2001; Butts. B.C.: 289; TL: BC, St. Elias Mts, Tats Lake, 770m
-
Boloria improba (Butler, 1877)
. See [maps]
Kääpiöhopeatäplä
Dusky-windged Fritillary
Dinky Fritillary
Dingy Arctic Fritillary
Dvärgpärlefjäril
- Argynnis improba Butler, 1877; Ent. mon. Mag. 13: 206; TL: "Winter Cove and Cambridge Bay" [NW Territories, Canada]
- Clossiana improba; [NACL], #4467; [BARU, #197]; [BOW]: pl. 13, f. 20; [NHM card]; [BRU2]: 47, pl. 33, f. 22-24; [RFEL]
- Boloria improba; [BNA]: 321, #245, pl. 5, 26, 60; [Opler]; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 33
- Boloria (Clossiana) improba; Pelham, 2008, J. Res. Lepid. 40: 301
![[improba-1.jpg]](_improba-1.jpg)
600x450 (~38Kb) Gohpascurro/Bjorkliden, 1100-1400m, Torne Lapmark, Sweden 9 June 1985, Photo © Lars Andersen
|
![[improba-2v.jpg]](_improba-2v.jpg)
800x583 (~75Kb) underside Gohpascurro/Bjorkliden, 1100-1400m, Torne Lapmark, Sweden 9 June 1985, Photo © Lars Andersen
|
![[improba-3u.jpg]](_improba-3u.jpg)
800x549 (~56Kb) upperside Gohpascurro/Bjorkliden, 1100-1400m, Torne Lapmark, Sweden 9 June 1985, Photo © Lars Andersen
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![[improba-4u.jpg]](_improba-4u.jpg)
800x709 (~76Kb) upperside Gohpascurro/Bjorkliden, 1100-1400m, Torne Lapmark, Sweden 9 June 1985, Photo © Lars Andersen
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|
Larva on
Bistorta vivipara,
Salix herbacea [BRU2], 47,
Polygonum viviparum,
Salix [BARU],
Salix ,
S. reticulata?,
(NW.Terr.) [BNA], 321
-
Boloria erda (Christoph, 1893)
-. See [maps]
Larva on
Vaccinium vitis-idaea [BRU2], 50
-
-
B. e. dulkeiti (Kurentzov, 1970) Magadan
- Brenthis dulkeiti Kurentzov, 1970; Butterflies of the USSR Far East: 83, 108; TL: Omsukchansky Mts., Magadan
-
B. e. kitoica (Belik, 1996) E.Sayan
- Clossiana erda kitoica Belik, 1996; Phegea 24 (4): 163; TL: Vostochnyy Sayan mtns
-
B. e. puella (Churkin, 1999) Transbaikalia (Barguzinsky Mts.)
- Clossiana erda puella Churkin, 1999; Atalanta 29 (1/4): 121; TL: Buryatia, Barguzin Mts., Nesterikha River, Kedrovoe Lake, 1900-2000m
-
Boloria alberta (Edwards, 1890)
Alberta, British Columbia. See [maps]
Alberta Fritillary
Alberta Alpine Fritillary
- Brenthis alberta banffensis; [NHM card]
- Brenthis alberta; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
- Clossiana alberta; [NACL], #4472; [BARU, #204]; [NHM card]; [BOW]: pl. 13, f. 17; [BRU2]: 50, pl. 35, f. 1-3
- Boloria alberta; [BNA]: 322, #246, pl. 26; [Opler]
- Boloria (Clossiana) alberta; Pelham, 2008, J. Res. Lepid. 40: 303
![[alberta-1fu.jpg]](_alberta-1fu.jpg)
1600x1200 (~262Kb) upperside female Canada, Mt. Tripoli, 18.7.1989, Photo © Norbert Kondla leg.
|
![[alberta-1fv.jpg]](_alberta-1fv.jpg)
1600x1200 (~248Kb) underside female Canada, Mt. Tripoli, 18.7.1989, Photo © Norbert Kondla leg.
|
![[alberta-1mu.jpg]](_alberta-1mu.jpg)
1600x1200 (~233Kb) upperside male Canada, Mt. Tripoli, 18.7.1989, Photo © Norbert Kondla leg.
|
![[alberta-1mv.jpg]](_alberta-1mv.jpg)
1600x1200 (~226Kb) underside male Canada, Mt. Tripoli, 18.7.1989, Photo © Norbert Kondla leg.
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|
Larva on
Dryas octopetala,
Vaccinium [BARU],
Dryas octopetala,
Dryas integrifolia [BRU2], 50
-
Boloria polaris (Boisduval, [1828])
Arctic Europe, Arctic Asia, Arctic America. See [maps]
Pohjanhopeatäplä
Polar Fritillary
Polaris Fritillary
Polarpärlefjäril
- Argynnis polaris Boisduval, [1828]; Eur. Lepid. Index Meth. 1: 15; TL: Nordkap Peninsula [Norway]
- Brenthis polaris ab. transversa Lingonblad, 1944; Notulae Ent. 24 (1-2) (1-2): 50; TL: Finland, Malla
- Brenthis polaris; Dyar, 1903, Bull. U.S. nat. Mus. 52: 16
- Clossiana polaris; [NACL], #4470; [BOW]: pl. 2, f. 1; [NHM card]; [BARU, #202]; [BRU2]: 48, pl. 33, f. 1-3; [RFEL]
- Boloria polaris; [BNA]: 323, #248, pl. 26; [Opler]; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
- Boloria (Clossiana) polaris; Pelham, 2008, J. Res. Lepid. 40: 302
![[polaris-1fu.jpg]](_polaris-1fu.jpg)
660x498 (~61Kb) upperside female FINLAND: Li: Utsjoki Puollamoaivi, 775:50, 8.7.1997, Photo © Markku Savela
|
Larva on
Cassiope tetragona,
Vaccinium uliginosum,
Dryas octopetala,
D. punctata [BRU2], 48
-
B. p. polaris
- ?Argynnis polaris var. homeyeri Wocke, 1875; in Homeyer, Dt. Nordpolarfart 2: [?? 403 not there]; TL: Grondland
- Argynnis chariclea var. groenlandica [Skinner], 1892; Ent. News 3 (3): 49, pl. 2, f. 9-10; TL: Greenland
- Argynnis polaris var. americana Strand, 1905; Rep. 2nd Norweg. Arctic Exp. (3): 10
- Clossiana alberta kurenzovi Wyatt, 1961; Zs. Wiener ent. Ges. 46: 98, pl. 5, f. 1-2; TL: "Berge von Tschukotka", NE.Siberia
- Clossiana polaris digna Churkin, 2001; Helios 2: 97, pl. 8, f. 4-6; TL: Wrangel Is., Neizvestnaya R.
- Boloria polaris digna; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
- Clossiana alberta kurenzovi; [BARU]; [NHM card]; [BRU2], 50
- Clossiana erda kurenzovi; [RFEL]
- Boloria erda kurenzovi; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
- Argynnis polaris v. americana; Strand, 1912, Ent. Zs. 25 (49): 253
- Clossiana polaris americana; [NHM card]
- Boloria polaris polaris; Pelham, 2008, J. Res. Lepid. 40: 302; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 34
-
B. p. stellata Masters, 1972 Manitoba
-
Boloria astarte (Doubleday, [1848])
Alaska, Washington, Montana. See [maps]
Astarte Fritillary
Arctic Ridge Fritillary
- Melitaea astarte Doubleday, [1848]; Gen. diurn. Lep. (2): pl. 53, f. 5; TL: Rock Lake, Alberta
Larva on
Saxifraga bronchialis [NAB]
-
Boloria tritonia (Böber, 1812)
-. See [maps]
Larva on
Saxifraga bronchialis Korshunov & Gorbunov, 1995, Butts. Asian Russia: 108
If you have corrections, comments or
information to add into these pages, just send mail to
Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.
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Life
Insecta
Lepidoptera
Ditrysia
Papilionoidea
Nymphalidae
Heliconiinae
![[sp-1.jpg]](sp-1.jpg)
700x504 (~51Kb) USA: Corral Pass region (47°00'15N 121°29'02W), Pierce Co., WA, 18.7.2000, Photo © Markku Savela
Cantabrian, Pyrenees, Alps, Carpathians, Caucasus, C.Asia, W.China. See [maps]
Shepherd's Fritillary
Perlada alpina
Dostojka pales
![[korla-1v.jpg]](korla-1v.jpg)
800x600 (~99Kb) underside An ancient moraine at 3400 m above sea level in the Tyshkan River headwaters, 45 km NEN of Zharkent, 20 km NEN of village Sarybel', altitude of 3400 m, mountain massif Tyshkantau, the Southern Dzhungarian Alatau mountain chain (NE part of Tien Shan), Zharkent District, Taldy-Kurgan Province, SE Kazakhstan. 26th of July 1994, Photo © Oleg Kosterin
W.Himalaya. See [maps]
Pyrenees, Alps, Fennoscandia. See [maps]
Tunturihopeatäplä
Mountain Fritillary
Napaea Fritillary
Fjällpärlefjäril
![[altaica-1mv.jpg]](altaica-1mv.jpg)
400x533 (~48Kb) underside male An alpine meadow, 2200 m above sea level, on the eastern spurs of the Katunskiy Mt. Range, the valley of the Argem (Direntai) stream, Central Altai Mts., West Siberia, Russia. 17th July, 1988 (on an Aster alpinum L. flower), Photo © Oleg Kosterin
![[altaica-2mfu.jpg]](altaica-2mfu.jpg)
652x500 (~54Kb) upperside male female A damp meadow on the Samakha Rivulet bank in the hollow Step' Samakha, 1600 m above sea level, at the junction of Argut and Koksu Rivers, Kosh-Agach District, Altai Republic, Central Altai Mts., West Siberia, Russia. 9th July, 1988 (on Bistorta bistorta), Photo © Oleg Kosterin
![[altaica-2mfvu.jpg]](altaica-2mfvu.jpg)
538x464 (~27Kb) upperside underside male female Highland dwarf birch (Betula rotundifolia) tundras on a ledge of the southerm principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets, 2400 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 10th July 1998, Photo © Oleg Kosterin
![[altaica-3mu.jpg]](altaica-3mu.jpg)
790x560 (~86Kb) upperside male a subalpine parkland of Pinus sibirica Du Tour on the Seminskii Pass through the Seminskii Mt. Range, at 1800 m above sea level. Ongudai District, Altai Republic, West Siberia, Russia. 9th July 1999 (on Betula nana rotundifolia), Photo © Oleg Kosterin
![[altaica-3mv.jpg]](altaica-3mv.jpg)
498x578 (~68Kb) underside male a subalpine parkland of Pinus sibirica Du Tour on the Seminskii Pass through the Seminskii Mt. Range, at 1800 m above sea level. Ongudai District, Altai Republic, West Siberia, Russia. 9th July 1999 (on Aster alpinus), Photo © Oleg Kosterin
Buryatia (Barguzin Mts.). See [maps]
Alaska - British Columbia, Arctic America. See [maps]
Altai, Sayan, Tuva (mountains), W.Mongolia. See [maps]
![[frigidalis-1fv.jpg]](frigidalis-1fv.jpg)
300x496 (~17Kb) underside female An alpine meadow vegetation at a brook flowing in a moss/sedge tundra in a cirque of the sources of a rivulet descending from the southerm principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets, 2650 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 16 July 1998 (on Rhodiola algida (Ledeb.) Fisch. & Mey.), Photo © Oleg Kosterin
![[frigidalis-1fvx.jpg]](frigidalis-1fvx.jpg)
960x588 (~107Kb) underside female A chionophylic alpine meadow in a cirque of one of the Chikty Rivulet sources, 2700 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 13th July 1998 (on Allium schoenoprasum), Photo © Oleg Kosterin
![[frigidalis-1mv.jpg]](frigidalis-1mv.jpg)
482x654 (~23Kb) underside male A chionophylic alpine meadow in a cirque of one of the Chikty Rivulet sources, 2700 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 13th July 1998 (on Allium schoenoprasum), Photo © Oleg Kosterin
![[frigidalis-1mvx.jpg]](frigidalis-1mvx.jpg)
864x712 (~99Kb) underside male A chionophylic alpine meadow in a cirque of one of the Chikty Rivulet sources, 2700 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 13th July 1998 (on Allium schoenoprasum), Photo © Oleg Kosterin
![[frigidalis-2mu.jpg]](frigidalis-2mu.jpg)
597x400 (~38Kb) upperside male A chionophylic alpine meadow in a cirque of one of the Chikty Rivulet sources, 2700 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West SIberia, Russia. 13th July 1998, Photo © Oleg Kosterin
Caucasus, Transcaucasia, Turkey. See [maps]
![[caucasica-1au.jpg]](caucasica-1au.jpg)
1016x908 (~323Kb) upperside Russia, Adygea Republic, Caucasian nature reserve, Lagonaki Plateau, Blyam Mt. E. slope, 12 July 2008, Photo © Oleg Kosterin
![[caucasica-1av.jpg]](caucasica-1av.jpg)
1242x874 (~308Kb) underside Russia, Adygea Republic, Caucasian nature reserve, Lagonaki Plateau, Blyam Mt. E. slope, 12 July 2008, Photo © Oleg Kosterin
![[caucasica-1b.jpg]](caucasica-1b.jpg)
908x851 (~177Kb) Russia, Adygea Republic, Caucasian nature reserve, Lagonaki Plateau, Blyam Mt. E. slope, 12 July 2008, Photo © Oleg Kosterin
W.Siberia, C.Siberia. See [maps]
Suohopeatäplä
Cranberry Fritillary
Allmän gulfläckig pärlemofjäril
Tranpärlefjäril
Dostojka akwilonaris
Amur, E.Yakutia?, Kamchatka, Sakhalin?. See [maps]
![[banghaasi-1v.jpg]](banghaasi-1v.jpg)
533x400 (~50Kb) underside On the local dwarf birch Betula divaricata Ledeb. An Empetrum/lichen tundra alternating with dwarf pine Pinus pumila (Pall.) Regel elfin wood in the delta of the Burgauli River, the southern coast of the Koni Peninsula, Ola District, Magadan Province, Russia. 18th July, 1989, Photo © Oleg Kosterin
![[roddi-1amu.jpg]](roddi-1amu.jpg)
598x526 (~60Kb) upperside male Boggy, with Pentaphylloides fruticosa bushes left bank of the Dzhazator River in its upper flow, between the falls of its right tributaries Chikty and Akbul, 2000 m above sea level, Kosh-Agach District, Altai Republic, West Siberia, Russia. 19th July 1998, Photo © Oleg Kosterin
![[roddi-1bmu.jpg]](roddi-1bmu.jpg)
732x500 (~79Kb) upperside male Boggy, with Pentaphylloides fruticosa bushes left bank of the Dzhazator River in its upper flow, between the falls of its right tributaries Chikty and Akbul, 2000 m above sea level, Kosh-Agach District, Altai Republic, West Siberia, Russia. 19th July 1998, Photo © Oleg Kosterin
![[roddi-1bmv.jpg]](roddi-1bmv.jpg)
400x515 (~34Kb) underside male Boggy, with Pentaphylloides fruticosa bushes left bank of the Dzhazator River in its upper flow, between the falls of its right tributaries Chikty and Akbul, 2000 m above sea level, Kosh-Agach District, Altai Republic, West Siberia, Russia. 19th July 1998 (on Tripleurospermum ambiguum), Photo © Oleg Kosterin
Balkans, Alps. See [maps]
Balkan Fritillary
. See [maps]
Rämehopeatäplä
Bog Fritillary
Ocellate Bog Fritillary
Svarttringlad pärlemorfjäril
Odonpärlefjäril
Randring-Perlmutterfalter
Dostojka eunomia
![[eunomia-1u.jpg]](eunomia-1u.jpg)
450x713 (~65Kb) upperside
![[eunomia-1v.jpg]](eunomia-1v.jpg)
427x675 (~66Kb) underside
![[eunomia-2u.jpg]](eunomia-2u.jpg)
419x352 (~33Kb) upperside
![[eunomia-2v.jpg]](eunomia-2v.jpg)
409x342 (~34Kb) underside
![[eunomia-3u.jpg]](eunomia-3u.jpg)
780x500 (~92Kb) upperside Russia, Moscow area, 15.6.2011 (36°25'E, 56°00'N), Photo © D. Smirnov
![[eunomia-3v.jpg]](eunomia-3v.jpg)
1009x577 (~114Kb) underside Russia, Moscow area, 17.6.2011 (36°25'E, 56°00'N), Photo © D. Smirnov
![[acidalia-1mv.jpg]](acidalia-1mv.jpg)
533x400 (~49Kb) underside male An alpine meadow on a cirque bottom at 2500 m above sea level in the Argem (Direntai) Stream headwaters, 15 km W of the village Dzhazatr on the eastern spurs of the Katunskiy Mt. Range, Kosh-Agach District, Central Altai Mts., West Siberia, Russia. 19th July, 1988, Photo © Oleg Kosterin
![[selene-1u.jpg]](selene-1u.jpg)
450x697 (~67Kb) upperside
![[selene-1v.jpg]](selene-1v.jpg)
458x713 (~67Kb) underside
![[selene-2u.jpg]](selene-2u.jpg)
425x374 (~35Kb) upperside
![[selene-2v.jpg]](selene-2v.jpg)
401x353 (~34Kb) underside
![[selene-3u.jpg]](selene-3u.jpg)
500x391 (~75Kb) upperside Haldon woods, Devon, UK 21/05/95, Photo © Simon Coombes
![[selene-3v.jpg]](selene-3v.jpg)
500x389 (~57Kb) underside Haldon woods, Devon, UK 21/05/95, Photo © Simon Coombes
![[selene-4v.jpg]](selene-4v.jpg)
837x1028 (~120Kb) underside Finland, Kuhmalahti, Mattilankylä, Setälänlahti, 23.6.2005, Photo © Harri Arkkio
![[selene-4vx.jpg]](selene-4vx.jpg)
1371x992 (~199Kb) underside Finland, Kuhmalahti, Mattilankylä, Setälänlahti, 23.6.2005, Photo © Harri Arkkio
![[selene-5fu.jpg]](selene-5fu.jpg)
852x639 (~123Kb) upperside female Russia, Moscow area (36°25'E, 56°00'N), 14.6.2009, Photo © D. Smirnov
![[selene-5mu.jpg]](selene-5mu.jpg)
711x624 (~103Kb) upperside male Russia, Moscow area, 14.6.2010 (36°25'E, 56°00'N), Photo © D. Smirnov
![[selene-5v.jpg]](selene-5v.jpg)
824x689 (~114Kb) underside Russia, Moscow area, 10.6.2010 (36°25'E, 56°00'N), Photo © D. Smirnov
![[selene-6mu.jpg]](selene-6mu.jpg)
918x694 (~122Kb) upperside male Russia, Moscow area, 3.7.2010 (36°25'E, 56°00'N), Photo © D. Smirnov
![[selene-6mv.jpg]](selene-6mv.jpg)
726x732 (~106Kb) underside male Russia, Moscow area, 3.7.2010 (36°25'E, 56°00'N), Photo © D. Smirnov
![[selene-7u.jpg]](selene-7u.jpg)
700x700 (~78Kb) upperside A grassy open larch forest on the Yabaganskii Pass (1400 m above sea level) through the Seminskii mountain range, the northern Altai Mts., West Siberia, Russia. 11th July 1999, Photo © Oleg Kosterin
![[selene-8u.jpg]](selene-8u.jpg)
916x545 (~97Kb) upperside Russia, Moscow area, 15.6.2011 (36°25'E, 56°00'N), Photo © D. Smirnov
![[tienschanica-1v.jpg]](tienschanica-1v.jpg)
400x533 (~49Kb) underside An alpine meadow on the right bank terrace of the fourth tributary of the Sargir River (the Usek River basin), altitude 3200 m, 47 km NEN of Zharkent, 21 km NEN of village Sarybel', the mountain massif Tyshkantau, the Southern Dzhungarian Alatau mountain chain (NE part of Tien Shan), Zharkent District, Taldy-Kurgan Province, SE Kazakhstan. 27th of July 1994, Photo © Oleg Kosterin
![[sibirica-1v.jpg]](sibirica-1v.jpg)
532x326 (~51Kb) underside A southern steppefied slope of the mountain on the Chuya River right bank between the villages Kurai and Chagan-Uzun, Altai Mts., West SIberia, Russia. 7th July 1988, Photo © Oleg Kosterin
![[oscarus-1mv.jpg]](oscarus-1mv.jpg)
488x400 (~38Kb) underside male A bushy meadow on the second right bank terrace of the Koen River at the Volchikha Rivulet just upstream of its fall into the Koen River near the village Nizhniy Koen, Iskitim District, Novosibirsk Province, West Siberia, Russia. 18th June, 1989, Photo © Oleg Kosterin
![[euphrosyne-1u.jpg]](euphrosyne-1u.jpg)
500x349 (~71Kb) upperside Luckett, Cornwall, UK 20/05/95, Photo © Simon Coombes
![[euphrosyne-1v.jpg]](euphrosyne-1v.jpg)
417x500 (~57Kb) underside Luckett, Cornwall, UK 20/05/95, Photo © Simon Coombes
![[euphrosyne-2l.jpg]](euphrosyne-2l.jpg)
533x400 (~50Kb) larva An open pine grove above the bluff of the valley of the Bol'shoi Elbash River, 5 km NW of the village Ust'-Chem, Iskitim dist., Novosibirsk region, West Siberia, 9th May 1995 (mature larva), Photo © Oleg Kosterin
![[euphrosyne-3v.jpg]](euphrosyne-3v.jpg)
600x832 (~62Kb) underside Finland, Loppi, Luutasuo 673:35, 2.6.1979, Photo © Pertti Pakkanen
![[euphrosyne-4fu.jpg]](euphrosyne-4fu.jpg)
900x745 (~146Kb) upperside female FINLAND: Li: Inari, Luttojoki, 760:54, 11.7.1998, Photo © Markku Savela
![[euphrosyne-5u.jpg]](euphrosyne-5u.jpg)
600x722 (~61Kb) upperside FINLAND, Ab: Suomusjärvi, 669:31, 15.6.1997, Photo © Markku Savela
![[euphrosyne-6.jpg]](euphrosyne-6.jpg)
900x1291 (~125Kb) Finland, Ab: Suomusjärvi (669:31), 6.6.2004, Photo © Markku Savela
![[euphrosyne-7mu.jpg]](euphrosyne-7mu.jpg)
1258x972 (~356Kb) upperside male An alpine meadow patch neighbouring with the most elevated larch wood outposts in the Kamdyt valley, a Tekelyu River right tributary which in turn is an Akkem River right tributary, the Katunskii Range northern principal slope, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Republic, West Siberia, Russia. 30th June 2007, Photo © Oleg Kosterin
![[euphrosyne-7mv.jpg]](euphrosyne-7mv.jpg)
861x1128 (~219Kb) underside male An alpine meadow patch neighbouring with the most elevated larch wood outposts in the Kamdyt valley, a Tekelyu River right tributary which in turn is an Akkem River right tributary, the Katunskii Range northern principal slope, Central [Russian] Altai Mts., Ust'-Koksa District, Altai Rapublic, West Siberia, Russia. 2nd July 2007, Photo © Oleg Kosterin
![[euphrosyne-8u.jpg]](euphrosyne-8u.jpg)
779x555 (~89Kb) upperside Russia, Moscow area, 11.6.2011 (38°52'E, 55°33'N), Photo © D. Smirnov
![[euphrosyne-8v.jpg]](euphrosyne-8v.jpg)
632x611 (~109Kb) underside Russia, Moscow area, 11.6.2011 (38°52'E, 55°33'N), Photo © D. Smirnov
![[euphrosyne-v.jpg]](euphrosyne-v.jpg)
800x631 (~57Kb) underside FINLAND: Ka: Virolahti, 671:53 (Reskopinsuo), 14.6.1997, Photo © Markku Savela
![[pallida-1fv.jpg]](pallida-1fv.jpg)
533x400 (~67Kb) underside female A dwarf birch (Betula rotundifolia Spach) montane tundra at 2200 m above sea level in the Argem (Direntai) Stream basin, 15 km W of the village Dzhazatr, on the eastern spurs of the Katunskiy Mt. Range, Kosh-Agach District, Central Altai Mts., West Siberia, Russia. 13th July, 1988, Photo © Oleg Kosterin
![[pallida-1mu.jpg]](pallida-1mu.jpg)
508x444 (~38Kb) upperside male A meadowy-tundrous SW slope of a ridge between the headwaters of the Chikty Rivulet, 2600 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 10th July 1998 (on Aster alpinus), Photo © Oleg Kosterin
![[pallida-1xmu.jpg]](pallida-1xmu.jpg)
600x600 (~71Kb) upperside male A meadowy-tundrous SW slope of a ridge between the headwaters of the Chikty Rivulet, 2600 m above sea level, the southern principle slope of the Yuzhno-Chuiskii mountain range, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 10th July 1998 (on Aster alpinus), Photo © Oleg Kosterin
![[angarensis-1fu.jpg]](angarensis-1fu.jpg)
533x400 (~45Kb) upperside female An open larch (Larix gmelinii (Rupr.) Rupr.)/birch (Betula pendula Roth. and B. davurica Pallas) forest above the left bluff of the Budyumkan River valley 5 km of its confluence with the Argun' River 6 km S of the village Uryupino, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 2nd August 1997, Photo © Oleg Kosterin
![[angarensis-1fv.jpg]](angarensis-1fv.jpg)
352x299 (~22Kb) underside female A road in a larch (Larix gmelinii (Rupr.) Rupr.) forest not far from the Argun' River left bank between the village Uryupino and the Gazimur River, Gazimurskozavodskoi District, E Chita Province, E Transbaikalia, Siberia, Russia. 31th July 1997, Photo © Oleg Kosterin
![[thore-1u.jpg]](thore-1u.jpg)
600x600 (~98Kb) upperside forest-tundra belt of the Koni Peninsula, S Magadan Region, Russia. 11th of July 1989, Photo © Oleg Kosterin
![[thore-1v.jpg]](thore-1v.jpg)
374x506 (~29Kb) underside Among the dead branches of the Siberian dwarf pine (Pinus pumila Ledeb.) A forest/tundra on the Koni Peninsula, S Magadan Region, Russia. 11th of July 1989, Photo © Oleg Kosterin
![[thore-u.jpg]](thore-u.jpg)
494x684 (~70Kb) upperside
![[thore-v.jpg]](thore-v.jpg)
470x733 (~69Kb) underside
![[hypercala-1fu.jpg]](hypercala-1fu.jpg)
600x600 (~102Kb) upperside female An open spruce forest, the Stolby Nature Reserve, Krasnoyarsk Province, 11th July 1995, Photo © Oleg Kosterin
![[hypercala-2fum.jpg]](hypercala-2fum.jpg)
784x1098 (~137Kb) upperside male female A grassy open larch forest on the Yabaganskii Pass (1400 m above sea level) through the Seminskii mountain range, the northern Altai Mts., West Siberia, Russia. 11th July 1999, Photo © Oleg Kosterin
![[hypercala-2fv.jpg]](hypercala-2fv.jpg)
800x800 (~145Kb) underside female A grassy open larch forest on the Yabaganskii Pass (1400 m above sea level) through the Seminskii mountain range, the northern Altai Mts., West Siberia, Russia. 11th July 1999, Photo © Oleg Kosterin
![[hypercala-2mv.jpg]](hypercala-2mv.jpg)
1088x786 (~121Kb) underside male A grassy open larch forest on the Yabaganskii Pass (1400 m above sea level) through the Seminskii mountain range, the northern Altai Mts., West Siberia, Russia. 11th July 1999, Photo © Oleg Kosterin
![[frigga-u.jpg]](frigga-u.jpg)
461x381 (~37Kb) upperside
![[frigga-v.jpg]](frigga-v.jpg)
447x373 (~36Kb) underside
![[alpestris-1fu.jpg]](alpestris-1fu.jpg)
872x600 (~56Kb) upperside female Highland dwarf birch tundras on of the southerm principal slope of the Yuzhno-Chuiskii mountain at the the Chikty Rivulet headwaters, 2400 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 15th July 1998, Photo © Oleg Kosterin
![[alpestris-1fux.jpg]](alpestris-1fux.jpg)
900x592 (~98Kb) upperside female A flowery alpine meadow in a valley of the rivulet descending from the southerm principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets, 2600 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 11th July 1998, Photo © Oleg Kosterin
![[alpestris-1fvx.jpg]](alpestris-1fvx.jpg)
800x800 (~121Kb) underside female A flowery alpine meadow in a valley of the rivulet descending from the southerm principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets, 2600 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 11th July 1998, Photo © Oleg Kosterin
![[alpestris-1mv.jpg]](alpestris-1mv.jpg)
475x500 (~33Kb) underside male An alpine meadow vegetation at a brook flowing in a moss/sedge tundra in a cirque of the sources of a rivulet descending from the southerm principle slope of the Yuzhno-Chuiskii mountain range between the Chikty and Akbul Rivulets, 2650 m above sea level, the upper Dzhazator River basin, Kosh-Agach District, Altai Republic, West Siberia, Russia. 16 July 1998, Photo © Oleg Kosterin
![[dia-1v.jpg]](dia-1v.jpg)
533x400 (~44Kb) underside A dwarf birch (Betula rotundifolia Spach) montane tundra at 2200 m above sea level in the Argem (Direntai) Stream basin, 15 km W of the village Dzhazatr, on the eastern spurs of the Katunskiy Mt. Range, Kosh-Agach District, Central Altai Mts., West Siberia, Russia. 129 - 15th, 130 - 12th July, 1988, Photo © Oleg Kosterin
![[dia-2l.jpg]](dia-2l.jpg)
598x400 (~44Kb) larva Central Europe 1990-2000, Photo © Michael Borsch
![[dia-2u.jpg]](dia-2u.jpg)
598x400 (~45Kb) upperside Central Europe 1990-2000, Photo © Michael Borsch
![[dia-3u.jpg]](dia-3u.jpg)
1100x768 (~335Kb) upperside Russia, West Siberia, Altaiskii Krai Province, Zmeinogorsk District, Savvushka village environs, the Lake Kolyvenskoe bank, a slope steppe. 9th September 2007, Photo © Oleg Kosterin
![[dia-u.jpg]](dia-u.jpg)
465x500 (~43Kb) upperside Axat, Pyrenees orientales, France 24/07/96, Photo © Simon Coombes
![[dia-v.jpg]](dia-v.jpg)
500x503 (~50Kb) underside Axat, Pyrenees orientales, France 24/07/96, Photo © Simon Coombes
![[titania-1a.jpg]](titania-1a.jpg)
571x653 (~100Kb) Russia, Moscow area (36°25'E, 56°00'N), 24.7.2009, Photo © D. Smirnov
![[titania-1v.jpg]](titania-1v.jpg)
604x640 (~97Kb) underside Russia, Moscow area (36°25'E, 56°00'N), 24.7.2009, Photo © D. Smirnov
![[staudingeri-1u.jpg]](staudingeri-1u.jpg)
700x510 (~58Kb) upperside A grassy open larch forest on the Yabaganskii Pass (1400 m above sea level) through the Seminskii mountain range, the northern Altai Mts., West Siberia, Russia. 11th July 1999, Photo © Oleg Kosterin
![[chariclea-1u.jpg]](chariclea-1u.jpg)
550x424 (~48Kb) upperside FINLAND: Li: Utsjoki Puollamoaivi, 775:50, 8.7.1997, Photo © Markku Savela
![[chariclea-1v.jpg]](chariclea-1v.jpg)
500x564 (~35Kb) underside FINLAND: Li: Utsjoki Ailigas, 775:50, 8.7.1997, Photo © Markku Savela
![[chariclea-2u.jpg]](chariclea-2u.jpg)
700x506 (~69Kb) upperside FINLAND: Li: Utsjoki Ailigas, 775:50, 6.7.1997, Photo © Markku Savela
![[helena-1fu.jpg]](helena-1fu.jpg)
802x949 (~107Kb) upperside female USA: Colorado, Clear Creek Co., Summit Lake, Mt. Evans, 23.7.2012 (39°35'52"N 105°38'23"W), Photo © Markku Savela
![[helena-2av.jpg]](helena-2av.jpg)
800x998 (~123Kb) underside USA: Colorado, Clear Creek Co., Summit Lake, Mt. Evans, 23.7.2012 (39°35'52"N 105°38'23"W), Photo © Markku Savela
![[helena-2bv.jpg]](helena-2bv.jpg)
800x608 (~84Kb) underside USA: Colorado, Clear Creek Co., Summit Lake, Mt. Evans, 23.7.2012 (39°35'52"N 105°38'23"W), Photo © Markku Savela
![[epithore-1uU.jpg]](epithore-1uU.jpg)
identification uncertain 600x539 (~44Kb) upperside USA: Near Crystal Mtn, (46°57'10N 121°28'27W), Pierce Co., WA, 27.7.1999, Photo © Markku Savela
![[epithore-1vU.jpg]](epithore-1vU.jpg)
identification uncertain 400x369 (~20Kb) underside USA: Near Crystal Mtn, (46°57'10N 121°28'27W), Pierce Co., WA, 27.7.1999, Photo © Markku Savela
![[epithore-2fu.jpg]](epithore-2fu.jpg)
1000x860 (~111Kb) upperside female USA: Washington, Kittitas Co., Tucquala Lake, 24.7.2011, Photo © Markku Savela
![[epithore-2v.jpg]](epithore-2v.jpg)
1000x837 (~96Kb) underside USA: Washington, Kittitas Co., Tucquala Lake, 24.7.2011, Photo © Markku Savela
![[epithore-3u.jpg]](epithore-3u.jpg)
1400x1046 (~252Kb) upperside USA: Washington, Skagit Co., on Sauk Mountain Rd (48°31'23N 121°36'22W), 15.8.2012, Photo © Markku Savela
![[epithore-3v.jpg]](epithore-3v.jpg)
1000x925 (~153Kb) underside USA: Washington, Skagit Co., on Sauk Mountain Rd (48°31'23N 121°36'22W), 15.8.2012, Photo © Markku Savela
![[epithore-4a.jpg]](epithore-4a.jpg)
1000x1249 (~153Kb) USA: Washington, Skagit Co., Sauk Mountain Trail Head & Trail, 15.8.2012, Photo © Markku Savela
![[epithore-4u.jpg]](epithore-4u.jpg)
1200x957 (~152Kb) upperside USA: Washington, Skagit Co., Sauk Mountain Trail Head & Trail, 15.8.2012, Photo © Markku Savela
![[improba-1.jpg]](improba-1.jpg)
600x450 (~38Kb) Gohpascurro/Bjorkliden, 1100-1400m, Torne Lapmark, Sweden 9 June 1985, Photo © Lars Andersen
![[improba-2v.jpg]](improba-2v.jpg)
800x583 (~75Kb) underside Gohpascurro/Bjorkliden, 1100-1400m, Torne Lapmark, Sweden 9 June 1985, Photo © Lars Andersen
![[improba-3u.jpg]](improba-3u.jpg)
800x549 (~56Kb) upperside Gohpascurro/Bjorkliden, 1100-1400m, Torne Lapmark, Sweden 9 June 1985, Photo © Lars Andersen
![[improba-4u.jpg]](improba-4u.jpg)
800x709 (~76Kb) upperside Gohpascurro/Bjorkliden, 1100-1400m, Torne Lapmark, Sweden 9 June 1985, Photo © Lars Andersen
![[alberta-1fu.jpg]](alberta-1fu.jpg)
1600x1200 (~262Kb) upperside female Canada, Mt. Tripoli, 18.7.1989, Photo © Norbert Kondla leg.
![[alberta-1fv.jpg]](alberta-1fv.jpg)
1600x1200 (~248Kb) underside female Canada, Mt. Tripoli, 18.7.1989, Photo © Norbert Kondla leg.
![[alberta-1mu.jpg]](alberta-1mu.jpg)
1600x1200 (~233Kb) upperside male Canada, Mt. Tripoli, 18.7.1989, Photo © Norbert Kondla leg.
![[alberta-1mv.jpg]](alberta-1mv.jpg)
1600x1200 (~226Kb) underside male Canada, Mt. Tripoli, 18.7.1989, Photo © Norbert Kondla leg.
![[polaris-1fu.jpg]](polaris-1fu.jpg)
660x498 (~61Kb) upperside female FINLAND: Li: Utsjoki Puollamoaivi, 775:50, 8.7.1997, Photo © Markku Savela