Theclinae
Nopsasiivet
Hairstreaks
Strong Blues
Theclinae (Lycaenidae) ; Clench, 1965, Mem. Amer. Ent. Soc. 19 : 331
Aphnaeinae (Lycaenidae) ; Clench, 1965, Mem. Amer. Ent. Soc. 19 : 357
Theclinae (Lycaenidae) ; Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6) : 381 ; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343 : 120; [NL4A] , 118; [AfrL ]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2 : 68; [RFEL]
Artopoetes Chapman, 1909
— 2 spp.
Laeosopis Rambur, 1858
— 1 sp.
Thecla Fabricius, 1807
— 3 spp.
1 sp.
Shirozua Sibatani & Ito, 1942
— 2 spp.
Cordelia Shirôzu & Yamamoto, 1956
— 3 spp.
Gonerilia Shirôzu & Yamamoto, 1956
— 5 spp.
Coreana Tutt, [1907]
— 1 sp.
Ussuriana Tutt, [1907]
— 7 spp.
Chaetoprocta de Nicéville in Marshall & de Nicéville, 1890
— 3 spp.
Protantigius Shirôzu & Yamamoto, 1956
— 1 sp.
Leucantigius Shirôzu & Murayama, 1951
— 1 sp.
Ravenna Shirôzu & Yamamoto, 1956
— 1 sp.
Antigius Sibatani & Ito, 1942
— 4 spp.
Wagimo Sibatani & Ito, 1942
— 6 spp.
Araragi Sibatani & Ito, 1942
— 3 spp.
Japonica Tutt, [1907]
— 5 spp.
Hypaurotis Scudder, 1876
— 1 sp.
Habrodais Scudder, 1876
— 2 spp.
Euaspa Moore, 1884
— 12 spp.
Howarthia Shirôzu & Yamamoto, 1956
— 13 spp.
Teratozephyrus Sibatani, 1946
— 10 spp.
Esakiozephyrus Shirôzu & Yamamoto, 1956
— 12 spp.
Neozephyrus Sibatani & Ito, 1942
— 7 spp.
Chrysozephyrus Shirôzu & Yamamoto, 1956
— 66 spp.
Iratsume Sibatani & Ito, 1942
— 1 sp.
Favonius Sibatani & Ito, 1942
— 14 spp.
1 sp.
Austrozephyrus Howarth, 1957
— 3 spp.
Shaanxiana Koiwaya, 1993
— 1 sp.
Yamamotozephyrus Saigusa, 1993
— 1 sp.
Nanlingozephyrus Wang & Pang, 1998
Yamatozephyrus Wang & Pang, 1998
Goldia Dubatolov & Korshunov, 1990
— 1 sp.
Thermozephyrus Inomata & Itagaki, 1986
— 1 sp.
Saigusaozephyrus Koiwaya, 1993
— 1 sp.
Sibataniozephyrus Inomata, 1986
— 3 spp.
Shizuyaozephyrus ?Koiwaya, ?2003
— 1 sp.
?Iozephyrus ?Wang Min, 2002
Proteuaspa ?Koiwaya, ?2003
— 1 sp.
Section Section Amblopala
Subtribe Oxylidina
Oxylidina (Theclini) ; [AfrL ]
Oxylidini (Theclinae) ;
Amblypodia Horsfield, [1829]
— 3 spp.
Iraota Moore, [1881]
Silverstreak Blues
— 6 spp.
Myrina Fabricius, 1807
Fig Tree Blues
— 5 spp.
Hemiolaus Aurivillius, 1922
— 4 spp.
Hypolycaena C. & R. Felder, 1862
Tits
Hairstreaks
— 46 spp.
Zeltus de Nicéville in Marshall & de Nicéville, 1890
— 1 sp.
Chliaria Moore, 1884
Tits
— 5 spp.
Leptomyrina Butler, 1898
— 9 spp.
Subtribe Deudorigini Doherty, 1886
Capys Hewitson, 1865
Protea Butteflies
— 17 spp.
Pamela Hemming, 1935
— 1 sp.
Section Section Neomyrina
Drina de Nicéville in Marshall & de Nicéville, 1890
— 7 spp.
Section Section Dapidodigma
Iolaus Hübner, [1819]
Sapphires
— 118 spp.
Stugeta Druce, 1891
— 7 spp.
Etesiolaus Stempffer & Bennett, 1959
— 3 spp.
Pratapa Moore, [1881]
Tufted Royals
— 5 spp.
Tajuria Moore, [1881]
Royals
— 32 spp.
Dacalana Moore, 1884
— 23 spp.
Maneca de Nicéville in Marshall & de Nicéville, 1890
— 1 sp.
Creon de Nicéville, [1896]
— 1 sp.
Bullis de Nicéville, 1897
— 3 spp.
Sukidion Druce, 1891
— 1 sp.
Purlisa Distant, 1881
— 1 sp.
Jacoona Distant, 1884
— 2 spp.
Neocheritra Distant, 1885
— 5 spp.
Manto de Nicéville, 1895
— 1 sp.
Mantoides Druce, 1896
— 1 sp.
Thrix Doherty, 1891
— 1 sp.
Charana de Nicéville in Marshall & de Nicéville, 1890
Mandarin Blues
— 2 spp.
Suasa de Nicéville in Marshall & de Nicéville, 1890
— 1 sp.
Section Section Britomartis
Paruparo Takanami, 1982
— 8 spp.
Matsutaroa Hayashi, Schröder & Treadaway, 1984
— 1 sp.
Rachana Eliot in Corbet & Pendlebury, 1978
— 5 spp.
Section Section Hypochrysops
Semanga Distant, 1884
— 2 spp.
Mota de Nicéville in Marshall & de Nicéville, 1890
— 1 sp.
Zinaspa de Nicéville in Marshall & de Nicéville, 1890
— 4 spp.
Surendra Moore, 1878
Acacia Blues
— 4 spp.
Tribe Zesiini
Zesiusinae Swinhoe, 1911; 153; TG : Zesius Hübner
Zesius Hübner, [1819]
— 2 spp.
Tribe Hypotheclini Eliot, 1973
Cheritra Moore, [1881]
Imperials
— 4 spp.
Ritra de Nicéville in Marshall & de Nicéville, 1890
— 1 sp.
Cheritrella de Nicéville, 1887
Imperials
— 1 sp.
Ticherra de Nicéville, 1887
— 1 sp.
Drupadia Moore, 1884
— 12 spp.
Ahmetia Özdikmen, 2008
— 1 sp.
Tribe Remelanini Eliot, 1973
Tribe Eumaeini Doubleday, 1847
Section Section Callophrys
Section Callophrys (Eumaeini) ; [NL4A] , 125
Section Section Brangas
Section Brangas (Eumaeini) ; [NL4A] , 118
Brangas Hübner, [1819]
— 11 spp.
Thaeides Johnson, Kruse & Kroenlein, 1997
— 5 spp.
Enos Johnson, Kruse & Kroenlein, 1997
— 5 spp.
Lamasina Robbins, 2002
— 6 spp.
Evenus Hübner, [1819]
— 12 spp.
Section Section Atlides
Section Atlides (Eumaeini) ; [NL4A] , 119
Section Section Micandra
Section Micandra (Eumaeini) ; [NL4A] , 120
Johnsonita Salazar & Constantino, 1995
— 7 spp.
Brevianta Johnson, Kruse & Kroenlein, 1997
— 9 spp.
Micandra Schatz, 1888
— 10 spp.
Timaeta Johnson, Kruse & Kroenlein, 1997
— 7 spp.
Rhamma Johnson, 1992
— 28 spp.
Phothecla Robbins in Lamas, 2004
— 2 spp.
Salazaria D'Abrera & Bálint, 2001
— 2 spp.
Temecla Robbins, 2004
— 6 spp.
Ipidecla Dyar, 1916
— 3 spp.
Penaincisalia Johnson, 1990
— 33 spp.
Podanotum Torres, Hall, Willmott & Johnson, 1996
— 8 spp.
Busbiina Robbins, 2004
— 1 sp.
Section Section Thereus
Section Thereus (Eumaeini) ; [NL4A] , 122
Section Section Satyrium
Section Satyrium (Eumaeini) ; [NL4A] , 124
Section Section Thestius
Section Thestius (Eumaeini) ; [NL4A] , 126
Section Section Allosmaitia
Section Allosmaitia (Eumaeini) ; [NL4A] , 127
Section Section Lamprospilus
Section Lamprospilus (Eumaeini) ; [NL4A] , 127
Section Section Strymon
Section Strymon (Eumaeini) ; [NL4A] , 130
Tmolus Hübner, [1819]
— 6 spp.
Nicolaea Johnson, 1993
— 26 spp.
Ministrymon Clench, 1961
— 20 spp.
Exorbaetta Johnson, Austin, Le Crom & Salazar, 1997
— 1 sp.
Gargina Robbins, 2004
— 8 spp.
Siderus Kaye, 1904
— 8 spp.
Theclopsis Godman & Salvin, [1887]
— 8 spp.
Ostrinotes Johnson, Austin, Le Crom & Salazar, 1997
— 9 spp.
Strephonota Johnson, Austin, Le Crom & Salazar, 1997
— 25 spp.
Section Section Panthiades
Section Panthiades (Eumaeini) ; [NL4A] , 134
Section Section Hypostrymon
Hypostrymon Clench, 1961
— 4 spp.
Marachina Robbins, 2004
— 3 spp.
Apuecla Robbins, 2004
— 3 spp.
Nesiostrymon Clench, 1964
— 8 spp.
Balintus D'Abrera, 2001
— 1 sp.
Aubergina Johnson, 1991
— 5 spp.
Terenthina Robbins, 2004
— 2 spp.
Iaspis Kaye, 1904
— 10 spp.
Celmia Johnson, 1991
— 6 spp.
Dicya Johnson, 1991
— 6 spp.
Trichonis Hewitson, 1865
— 2 spp.
Section Section Erora
Section Erora (Eumaeini) ; [NL4A] , 136
Tribe Catapaecilmatini Eliot, 1973
Catapaecilmini[sic?] ; [MRS] , 638
Section Section Catapaecilma
Tribe Tomarini Eliot, 1973
Tribe Aphnaeini
Aphnaeini (Theclinae) ; Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6) : 381 , 435
Section Pseudaletis (Aphnaeini) ; Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6) : 436
Section Aphnaeus (Aphnaeini) ; Eliot, 1973, Bull. Br. Mus. nat. Hist. (Ent.) 28 (6) : 436
Aphnaeini (Theclinae) ; Heath, 1997, Metamorphosis Occ. Suppl. 2 : 9; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343 : 130; [AfrL ]
Aphnaeinae (Lycaenidae) ; Korb & Bolshakov, 2011, Eversmannia Suppl. 2 : 68; Boyle, Kaliszewska, Espeland, Suderman, Fleming, Heath & Pierce, 2015, Syst. Ent. 40 : 177
Phasis Hübner, [1819]
— 4 spp.
Tylopaedia Tite & Dickson, 1973
— 1 sp.
Argyraspodes Tite & Dickson, 1973
— 1 sp.
Aloeides Hübner, [1819]
— 58 spp.
Erikssonia Trimen, 1891
— 4 spp.
Trimenia Tite & Dickson, 1973
— 5 spp.
Aphnaeus Hübner, [1819]
Silver Spots
— 35 spp.
Axiocerses Hübner, [1819]
Scarlets
— 19 spp.
Zeritis Boisduval, [1836]
— 6 spp.
Crudaria Wallengren, 1875
— 3 spp.
Cigaritis Donzel, 1847
Leopard Butterflies
Silverlines
Barred Blues
— 72 spp.
Chloroselas Butler, [1886]
Gems
— 13 spp.
Lipaphnaeus Aurivillius, 1916
— 4 spp.
Chrysoritis Butler, [1898]
— 42 spp.
Vansomerenia Heath, 1997
— 1 sp.
Cesa Seven, 1997
— 1 sp.
Pseudaletis Druce, 1888
— 29 spp.
363x500 (~25Kb) Venezuela, Merida and the Sierra Nevada National Park, July 2002, Photo © Simon Coombes
The exact identification of this species is still unknown,
but tentatively assumed to belong into this group.
Unknown or unplaced taxa
7.4.2019 (2571)
Some related literature:
Boyle, Kaliszewska, Espeland, Suderman, Fleming, Heath & Pierce, 2015
Phylogeny of the Aphnaeinae: myrmecophilous African butterflies with carnivorous and herbivorous life histories
Syst. Ent. 40 : 169-182
[MRS]; Chou, 1994
Monografia Rhopalocerorum Sinensium 1-2: 1-854
Eliot, 1973
The higher classification of the Lycaenidae (Lepidoptera): a tentative Arrangement
Bull. Br. Mus. nat. Hist. (Ent.) 28 (6)
: 373 -505 (162fgs), pl. 1 -6
Clench in Fox, Lindsey, Clench & Miller, 1965
The Butterflies of Liberia
Mem. Amer. Ent. Soc. 19
: 1 -438
Heath, 1997
A review of African genera of the tribe Aphnaeini (Lepidoptera: Lycaenidae)
Metamorphosis Occ. Suppl. 2
: 1-60
Johnson, 1993
A New Neotropical Genus of Branded Angulopine-like Hairstreak Butterflies (Lepidoptera, Lycaenidae, Theclinae)
Rep. Mus. Nat. Hist. Univ. Wisc. (Stevens Point) 40
: 1-7
Korb & Bolshakov, 2011
[A catalogue of butterflies (Lepidoptera: Papilioformes) of the former USSR. Second edition, reformatted and updated] (in Russian)
Eversmannia Suppl. 2
: 1-123
[NL4A]; Lamas, 2004
Checklist: Part 4A, Hesperioidea - Papilionoidea
Atlas of Neotropical Lepidoptera
[RFEL]; Leley, 2016
Annotated catalogue of the insects of Russian Far East. Volume II. Lepidoptera
Cat. ins. Russian Far East. 2
: 1-812
Strand, 1916
Abbildungen und Beschreibungen neuer und wenig bekannter Lepidoptera aus der Sammlung W. Niepelt
Lepidoptera Niepeltiana (2)
: 1 -26, pl. 13 -17
Vane-Wright & de Jong, 2003
The butterflies of Sulawesi: annotated checklist for a critical island faunda
Zool. Verh. Leiden 343 : 3-268, pl. 1-16
If you have corrections, comments or
information to add into these pages, just send mail to
Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.