Life Insecta Lepidoptera Ditrysia Papilionoidea Lycaenidae Polyommatinae

600x400 (~104Kb) upperside India: Subansiri, 2.11.2005, Photo © Kedar Tokekar	600x400 (~101Kb) upperside India: Subansiri, 2.11.2005, Photo © Kedar Tokekar	600x400 (~79Kb) underside India: Subansiri, 2.11.2005, Photo © Kedar Tokekar
600x400 (~49Kb) India: Subansiri, 4.11.2005, Photo © Kedar Tokekar

The exact identification of these species is still unknown, but tentatively assumed to belong into this group.

• Acytolepis puspa (Horsfield, [1828]) Peninsular India, Himalayas, Assam, Ceylon, Burma, Philippines, Nicobars, New Guinea. See [maps] Common Hedge Blue

  • Polyommatus puspa Horsfield, [1828]; Descr. Cat. lep. Ins. Mus. East India Coy (1): 67; TL: Java
  • Cyaniris puspa var. watasei Matsumura, 1915; Ent. Mag. Kyoto 1 (2): 66, 68

  • Acytolepis puspa, Lepiforum [LEPIFORUM]

  • Cyaniris puspa; Druce, 1895, Proc. zool. Soc. Lond. 1895 (3): 572; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 284
  • Lycaenopsis puspa; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 437; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 24; [BIR], 274
  • Celastrina puspa; [BOW]: pl. 175, f. 20, 21
  • Acytolepis puspa; [MRS], 679; [NHM card]; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 161

  600x400 (~59Kb) underside India: Subansiri, 31.10.2005, Photo © Kedar Tokekar

  600x400 (~75Kb) underside India: Subansiri, 4.11.2005, Photo © Kedar Tokekar

  Larva on Glochidion fortunei, Rhododenron spp., Distylium racemosum, Rosa multiflora, Rosa wichuraina, R. centifolia, Prunus zippeliana, Glochidion obovatum, Myrica rubra, Quercus phillyraeoides, Celtis sinensis, Astilbe thunbergii [MRS], Schleichera oleosa, Hiptage benghalensis, Xylia dolabriformis [BMP], Viburnum , Combretaceae , Cycadaceae , Shorea , Enkianthus , Lyonia , Rhododendron , Vaccinium , Antidesma , Breynia , Bridelia , Glochidion , Calliandra , Cylista , Paracalyx , Pellophorum , Xylia , Quercus , Distylum , Hiptage , Myrica , Aruncus , Chaenomeles , Malus , Photinia , Prunus , Rosa , Spiraea , Dimocarpus , Euphoria , Litchi , Sapindus , Schleichera , Astilbe , Celtis Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 161

  • A. p. puspa

    • Lycaenopsis puspa puspa; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 26
  • A. p. cagaya (C. & R. Felder, [1865]) Philippines (Luzon)

    • Lycaena cagaya C. & R. Felder, [1865]; Reise Fregatte Novara, Bd 2 (Abth. 2) (2): 278, pl. 34, f. 11-13
      • = Lycaenopsis puspa ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 438
    • Cyaniris puspa ottonis Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 287; TL: Palawan

    • Lycaenopsis puspa ottonis; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25
    • Acytolepis puspa ottonis; [NHM card]
    • Lycaenopsis puspa cagaya; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25
    • Acytolepis puspa cagaya; [NHM card]
  • A. p. lambi (Distant, 1882) Siam, Vietnam, Tenasserim

    • Polyommatus lambi Distant, 1882; Ann. Mag. nat. Hist. (5) 10 (57): 245; TL: Malay Peninsula
      • = Lycaenopsis puspa ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 438
    • Cyaniris imperatrix Butler, 1900; Ann. Mag. nat. Hist. (7) 5 (29): 444; TL: Thailand
      • = Lycaenopsis puspa ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 438

    • Acytolepis puspa lambi, Butterflies in Indo-China [Yutaka Inayoshi]

    • Lycaenopsis puspa imperatrix; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25
    • Acytolepis puspa imperatrix; [NHM card]
    • Lycaenopsis puspa lambi; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25
    • Acytolepis puspa lambi; [NHM card]; [BMP]: 240, pl. 35, f. 44-46
  • A. p. puspinus (Kheil, 1884) Nias

    • puspa puspinus (Kheil, 1884) (Plebeius); Rhopalocera Ins. Nias: 30, f. 37-39; TL: Nias
      • = Lycaenopsis puspa ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 438

    • Cyaniris puspinus; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 287
    • Lycaenopsis puspa puspinus; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 26
    • Acytolepis puspa puspinus; [NHM card]
  • A. p. kuehni (Röber, 1886) Sulawesi

    • Plebeius kühni Röber, 1886; Corr.-Bl. Ent. Ver. Iris 1 (3): 60, pl. 4, f. 29 (partim); TL: E.Sulawesi
      • = Lycaenopsis puspa ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 438
    • Lycaenopsis puspa martini Fruhstorfer, 1922; in Seitz, Gross-Schmett. Erde 9: 870; TL: Sulawesi
      • = Acytolepis puspa kuehni ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 54
    • Lycaenopsis puspa hyllus Fruhstorfer, 1922; in Seitz, Gross-Schmett. Erde 9: 870; TL: Banggai
      • = Acytolepis puspa kuehni ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 54
    • Cyanirus[sic] kühni kalawari Ribbe, 1926; Ent. Mitt. 15 (1): 89; TL: Sulawesi, Kalawara
      • = Acytolepis puspa kuehni ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 54
    • Cyaniris kühni bangkaiensis Ribbe, 1926; Ent. Mitt. 15 (1): 89; TL: Banggai
      • = Acytolepis puspa kuehni ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 54

    • Cyaniris kuhni kalawari; [NHM card]
    • Cyaniris kuehni bangkaiensis; [NHM card]
    • Acytolepis puspa hyllus; [NHM card]
    • Lycaenopsis puspa martini; [NHM card]
    • Lycaenopsis puspa kuehni; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 438
    • Cyaniris kuhni[sic, recte kuehni]; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 287; [NHM card]
    • Lycaenopsis puspa kühni; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 26
    • Acytolepis puspa kuehni; Toxopeus, 1928, Tijdschr. Ent. 71: 231; Eliot & Kawazoé, 1983, Butts. Lycaenopsis group: 180; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 162
    • Acytolepis puspa kühni; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 54, f. 23-24
  • A. p. cyanescens (de Nicéville, 1890) Nicobar Is.

    • Cyaniris cyanescens de Nicéville, 1890; Butts India Burmah Ceylon 3: 103
      • = Lycaenopsis puspa ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 438

    • Lycaenopsis puspa cyanescens; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25
    • Acytolepis puspa cyanescens; [NHM card]
  • A. p. prominens (de Nicéville, 1890)

    • Cyaniris cyanescens prominens de Nicéville, 1890; Butts India Burmah Ceylon 3: 103

    • Lycaenopsis puspa prominens; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25
    • Acytolepis puspa prominans[sic, recte prominens]; [NHM card]
  • A. p. dammae (Heron, 1894) Timor, Damar, Wetar

    • Cyaniris dammae Heron, 1894; Ann. Mag. nat. Hist. (6) 14 (80): 104; TL: Damma I.
      • = Lycaenopsis puspa ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 438

    • Cyaniris puspa damae[sic, recte dammae]; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 286
    • Lycaenopsis puspa dammae; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 26
    • Acytolepis puspa dammae; [BAUR], 383; [NHM card]
  • A. p. timorensis (Butler, 1900) Timor

    • Cyaniris timorensis Butler, 1900; Ann. Mag. nat. Hist. (7) 5 (29): 443; TL: Timor
      • = Lycaenopsis puspa ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 438

    • Lycaenopsis puspa timorensis; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 26
    • Cyaniris puspa timorensis; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 286 (note)
    • Acytolepis puspa timorensis; [NHM card]
  • A. p. splendens (Butler, 1900)

    • Cyaniris splendens Butler, 1900; Ann. Mag. nat. Hist. (7) 5 (29): 444; TL: Larut Hills, Perak
      • = Lycaenopsis puspa ; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 438

    • Acytolepis puspa splendens; [NHM card]
  • A. p. myla (Fruhstorfer, 1909) Formosa

    • Cyaniris puspa myla Fruhstorfer, 1909; Ent. Zs. 23 (25): 116; TL: Formosa
    • Cyaniris postimacula Matsumura, 1929; Insecta Matsumurana 3 (2/3): 104; TL: Formosa
      • = Acytolepis puspa myla ; [NHM card]
    • myla (Matsumura, 1929) (Cyaniris); Insecta Matsumurana 3 (2/3): 104 (preocc. Fruhstorfer)
    • Celastrina puspa formosana Bethune-Baker, 1914; Ent. Mitt. 3 (4): 127; TL: Formosa
      • = Acytolepis puspa myla ; [NHM card]

    • Cyaniris puspa myla; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 284
    • Lycaenopsis puspa myla; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 24
    • Acytolepis puspa myla; [NHM card]; [MRS], 679

    Larva on Glochidion acuminatum, G. lanceolatum, G. rubrum, Bridelia balansae, Euphoria longan, Sapindus mukorossi, Vaccinium oldhamii, V. samllii, Enkyathe perulatus, Lyonia ovalifolia, Viburnum dilatatum, Prunus padus, Chaenomeles speciosa, Malus domestica, Spiraea cantoniensis, Aruncus dioicus, Astilbe thunbergii [MRS]

  • A. p. hermagoras (Fruhstorfer, 1910) Hainan

    • Cyaniris puspa hermagoras Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 284; TL: Hainan

    • Lycaenopsis puspa hermagoras; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25
    • Acytolepis puspa hermagoras; [NHM card]; [MRS], 679
  • A. p. sania (Fruhstorfer, 1910)

    • Cyaniris puspa sania Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 285

    • Lycaenopsis puspa sania; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 26
    • Acytolepis puspa sania; [NHM card]
  • A. p. gisca (Fruhstorfer, 1910) NE.Afghanistan - Burma, Thailand, Cambodia, Laos, Vietnam, S.China, Yunnan

    • Cyaniris puspa gisca Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 285; TL: Sikkim
    • Lycaenopsis puspa gisca f. artena Fruhstorfer, 1917; Archiv Naturg. 82 A (1): 25; TL: India
    • Lycaenopsis puspa telis Fruhstorfer, 1917; Archiv Naturg. 82 A (1): 25; TL: Andaman
    • Celastrina puspa barneyi Corbet, 1941; Hongkong Nat. 10 (3/4): 149; TL: Hong Kong

    • Acytolepis puspa gisca, Butterflies in Indo-China [Yutaka Inayoshi]

    • Acytolepis puspa artena; [NHM card]
    • Acytolepis puspa barneyi; [NHM card]
    • Acytolepis puspa telis; [NHM card]
    • Lycaenopsis puspa gisca; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25
    • Acytolepis puspa gisca; [MRS], 679; [NHM card]; Huang, 2003, Neue Ent. Nachr. 55: 106 (note)

    848x752 (~135Kb) underside male Cambodia, Ratanakiri Province, Virachey National Park biffer zone, Yorn village environs, Yak Kae Rivulet, 4th June 2013, Photo © Oleg Kosterin

  • A. p. hermione (Fruhstorfer, 1905) Bawean

    • Cyaniris puspa hermione Fruhstorfer, 1905; Soc. Ent. 20 (1): 2; TL: Bawean

    • Cyaniris puspa hermione; Fruhstorfer, 1905, Ent. Zs. 19 (12): 64, f. 10
    • Lycaenopsis puspa hermione; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 26
    • Acytolepis puspa hermione; [NHM card]
  • A. p. contilia (Fruhstorfer, 1910) Sumbawa

    • Cyaniris puspa contilia Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 285; TL: Sumbawa, Sumba

    • Lycaenopsis puspa contilia; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 26
    • Acytolepis puspa contilia; [NHM card]
  • A. p. bazilana (Fruhstorfer, 1910) Philippines (Mindoro, ...?)

    • Cyaniris puspa bazilana Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 286; TL: Bazilan
    • Cyaniris puspa georgi Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 286; TL: Camiguin de Mindanao
    • Lycaenopsis puspa sabis Fruhstorfer, 1917; Archiv Naturg. 82 A (1): 25

    • Acytolepis puspa sabis; [NHM card]
    • Lycaenopsis puspa georgi; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25
    • Acytolepis puspa georgi; [NHM card]
    • Lycaenopsis puspa bazilana; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25
  • A. p. calata (Fruhstorfer, 1910) Flores

    • Cyaniris puspa calata Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 286; TL: Flores

    • Lycaenopsis puspa calata; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 26
    • ?Udara placidula calata;
    • Acytolepis puspa calata; [NHM card]
  • A. p. darmis (Fruhstorfer, 1917) Kai, Aru?

    • Lycaenopsis puspa darmis Fruhstorfer, 1917; Archiv Naturg. 82 A (1): 27; TL: Key I.

    • Acytolepis puspa darmis; [BAUR], 383 (text)
  • A. p. mygdonia (Fruhstorfer, 1917) Sumatra, Borneo

    • Lycaenopsis puspa mygdonia Fruhstorfer, 1917; Archiv Naturg. 82 A (1): 26; TL: N.Sumatra; Borneo

    • Acytolepis puspa mygdonia; [NHM card]
  • A. p. roberi (Fruhstorfer, 1917) Goram

    • Lycaenopsis puspa röberi Fruhstorfer, 1917; Archiv Naturg. 82 A (1): 27; TL: Goram

    • Acytolepis puspa roberi; [BAUR], 383 (text); [NHM card]
  • A. p. volumina (Fruhstorfer, 1922)

    • Lycaenopsis volumina Fruhstorfer, 1922; in Seitz, Gross-Schmett. Erde 9: 869, Lfg. (302)

    • Acytolepis puspa volumina; [NHM card]
  • A. p. tymbria (Fruhstorfer, 1922)

    • puspa tymbria (Fruhstorfer, 1922) (Lycaenopsis); in Seitz, Gross-Schmett. Erde 9: 870

    • Acytolepis puspa tymbria; [NHM card]
  • A. p. deronda (Fruhstorfer, 1922) Sula Is.

    • Lycaenopsis puspa deronda Fruhstorfer, 1922; in Seitz, Gross-Schmett. Erde 9: 871; TL: Sula Is
    • Lycaenopsis puspa var. Chapman, 1909; Proc. zool. Soc. Lond. 1909 (2): 438

    • Acytolepis puspa deronda; [NHM card]; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 55; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 162
  • A. p. vandeldeni Toxopeus, 1929 Sumatra

    • Acytolepis puspa vandeldeni Toxopeus, 1929; Tijdschr. Ent. 72 (3/4): 207; TL: Pulu Weh

    • Acytolepis puspa vandeldeni; [NHM card]
  • A. p. felderi Toxopeus, 1927 Ceylon

    • Acytolepis puspa felderi Toxopeus, 1927; Tijdschr. Ent. 70 (3/4): 294

    • Acytolepis puspa felderi; [NHM card]
  • A. p. ishigakiana (Matsumura, 1929)

    • Cyaniris ishigakiana Matsumura, 1929; Insecta Matsumurana 3 (2/3): 104; TL: Okinawa

    • Celastrina ishigakiana; [NHM card]
  • A. p. umenonis (Matsumura, 1931) Japan

    • Lycaenopsis (Cyaniris) puspa f. umenonis Matsumura, 1931; Insecta Matsumurana 6 (1-2): 44

    • Celastrina puspa umenonis; Kudrna, 1974, Atalanta 5: 115
    • Acytolepis puspa umenonis; [NHM card]
  • A. p. haani (Kalis, 1938) Kangean I.

    • puspa haani (Kalis, 1938) (Lycaenopsis); Ent. Rdsch. 55: 198; TL: Kangean I.

    • Acytolepis puspa haani; [NHM card]
  • A. p. siporana (Riley, [1945]) Mentawi Is.

    • puspa siporana (Riley, [1945]) (Celastrina); Trans. R. ent. Soc. Lond. 94 (2): 256

    • Acytolepis puspa siporana; [NHM card]
• Acytolepis armenta (Fruhstorfer, 1910) Borneo. See [maps]

  • Cyaniris armenta Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 287; TL: Borneo; Lawas

  • Cyaniris armenta; [NHM card]
• Acytolepis lilacea (Hampson, 1889) Nilgiris, Palnis, Coorg. See [maps] Hampson's Hedge Blue

  • Cyaniris puspa var. lilacea Hampson, 1889; J. asiat. Soc. Bengal 57 (4): 356; TL: Nilgiris and Nellyampathy Hills, Cochin

  • Lycaenopsis lilacea; [BIR], 274
  • A. l. lilacea

    • Cyaniris puspa lilacina[sic, recte lilacea?]; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 285
    • Lycaenopsis puspa lilacea; ; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 25
    • Acytolepis puspa lilacea; [NHM card]
  • A. l. moorei (Toxopeus, 1926) Ceylon

    • Lycaenopsis moorei Toxopeus, 1926; Treubia 8 (3-4): 365

    • Acytolepis puspa moorei; [NHM card]
  • A. l. indochinensis Eliot & Kawazoé, 1983 Burma, Thailan, Laos

    • Acytolepis lilacea indochinensis Eliot & Kawazoé, 1983; Butts. Lycaenopsis group: 187, f. 135, 268, 284; TL: Vang Vien, Laos

    • Acytolepis lilacea indochinensis, Butterflies in Indo-China [Yutaka Inayoshi]

    • Acytolepis puspa indochinensis; [NHM card]
• Acytolepis ripte (Druce, 1895) Borneo. See [maps]

  • Cyaniris ripte Druce, 1895; Proc. zool. Soc. Lond. 1895 (3): 574, pl. 32, f. 11; TL: Labuan

  • Cyaniris ripte; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 301
  • ?Lycaenopsis rifte[sic, recte ripte?]; [NHM card]
  • Cyaniris ripte; [NHM card]
  • Lycaenopsis ripte; Chapman, 1909, Proc. zool. Soc. Lond. 1909 (2): 434; Fruhstorfer, 1917, Archiv Naturg. 82 A (1): 33
• Acytolepis najara (Fruhstorfer, 1910) Sulawesi. See [maps]

  • Cyaniris najara Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 287 (partim, ♂); TL: Sulawesi, Bua-Krang, 5000ft

  • Cyaniris najara; [NHM card]
  • Acytolepis najara; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 55; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 162
• Acytolepis samanga (Fruhstorfer, 1910) Samanga

  • Cyaniris samanga Fruhstorfer, 1910; Stettin ent. Ztg. 71 (2): 286; TL: Samangi, Bua
  • Lycaenopsis akasa parakasa Fruhstorfer, 1922; in Seitz, Gross-Schmett. Erde 9: 862; TL: Sulawesi, Minahassa
    • = ; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 55

  • Celastrina samanga; Eliot, 1956, Bull. Raffles Mus. 27: 35, f. 2
  • Acytolepis puspa samanga; Toxopeus, 1928, Tijdschr. Ent. 71: 231 (partim, female); [NHM card]
  • Acytolepis samanga; Cassidy, 1995, Trans. lepid. Soc. Japan 46 (2): 55; Vane-Wright & de Jong, 2003, Zool. Verh. Leiden 343: 162

  Misidentified/misapplied:

  • Cyaniris akasa; Pagenstecher, 1897, Abh. Senckenb. naturf. Ges. 23: 415
  • Cyaniris najara; Fruhstorfer, 1910, Stettin ent. Ztg. 71 (2): 287 (partim, 'female' recte male, nec male)
  • Celastrina lavendularis lyce; Toxopeus, 1928, Tijdschr. Ent. 71: 229 (partim, 'female' recte male, nec male)

25.12.2018 (6)

If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.