Bertula abjudicalis Walker, [1859]
Ceylon. See [maps ]
Bertula albipunctata Wileman, 1915 Formosa
Bertula albipunctata Wileman, 1915; 35; TL : Formosa, Kanshirei
Bertula albolinealis (Leech, 1900)
Bleptina albolinealis ; [POOLE]
Bertula albolinealis ; [MOB17] , 26 (implied)
Bertula albonotata (Bethune-Baker, 1908)
New Guinea. See [maps ]
Bleptina albonotata Bethune-Baker, 1908; 208; TL : British New Guinea, Aroa River
Bleptina albonotata ; [POOLE]
Bertula albonotata ; [MOB17] , 26 (implied)
Bertula albovenata (Leech, 1900)
Bleptina albovenata ; [POOLE]
Bertula albovenata ; [MOB17] , 26 (implied)
Bertula alpheusalis Walker, [1859]
NE.Himalaya, Taiwan, Peninsular Malaysia, Sumatra, Borneo. See [maps ]
Bertula ambigua (Leech, 1900)
Bertula angulifascia Holloway, 2008
Borneo, Peninsular Malaysia. See [maps ]
Bertula angulifascia Holloway, 2008; Moths of Borneo 17 : 45, f. 100, pl. 1; TL : Sarawak, Gunong Mulu Nat. Park, R.G.S. Exped.
Bertula atrirena Hampson, 1929
Philippines. See [maps ]
Bertula atrirena Hampson, 1929; 396; TL : Philippines, Luzon, Mt. Banahao
Bertula bidentata (Wileman, 1915) Formosa
Bleptina bidentata Wileman, 1915; 14; TL : Formosa, Arizan
Bertula bifurcana (Bethune-Baker, 1908)
New Guinea. See [maps ]
Bleptina bifurcana Bethune-Baker, 1908; 207; TL : British New Guinea, Aroa River
Bertula bifurcata Holloway, 2008
Borneo. See [maps ]
Bertula bifurcata Holloway, 2008; Moths of Borneo 17 : 30, f. 60, pl. 1; TL : Sandakan [Sabah]
Bertula bistrigata (Staudinger, 1888)
Japan, Korea, Amur, Ussuri. See [maps ]
Zanclognatha bistrigata Staudinger, 1888; 276; TL : Vladivostok
Bocana spacoalis chosenana Bryk, 1949; 150, pl. 6, f. 12; TL : Korea, Shuotsu
Bertula bornedescripta Holloway, 2008
Borneo. See [maps ]
Bertula bornedescripta Holloway, 2008; Moths of Borneo 17 : 47, f. 83, pl. 1; TL : Sarawak, Gunong Mulu Nat. Park, R.G.S. Exped.
Bertula caliginosa (Wileman, 1911)
Japan. See [maps ]
Bleptina caliginosa Wileman, 1911; 253; TL : Japan, Hakodate, Ishi Kaeva
Bleptina caliginosa ; [POOLE]
Bertula caliginosa ; [MOB17] , 26 (implied)
Bertula carta (Swinhoe, 1902)
Borneo, Peninsular Malaysia. See [maps ]
Bertula centralis (Wileman, 1915) Formosa
Bleptina centralis Wileman, 1915; 13; TL : Formosa, Kanshirei
Bertula contingens Walker, [1859]
Ceylon. See [maps ]
Bertula contingens Walker, [1859]; 245; TL : Ceylon
Bertula crucialis (Felder & Rogenhofer, 1874)
Sundaland. See [maps ]
Foveades crucialis ; [POOLE]
Bertula crucialis ; [MOB17] , 44, f. 99, pl. 1
Bertula curvilinea (Leech, 1900)
Bleptina curvilinea ; [POOLE]
Bertula curvilinea ; [MOB17] , 26 (implied)
Bertula delosticha (Swinhoe, 1906)
Sumatra, Borneo, Java. See [maps ]
Bleptina delosticha Swinhoe, 1906; 552; TL : W.Sumatra, Padang
Foveades delosticha ; [POOLE]
Bertula delosticha ; [MOB17] , 44, f. 43, 96, 127, pl. 1
Bertula dentilinea (Hampson, 1895)
Sikkim. See [maps ]
Bleptina (Bertula) dentilinea Hampson, 1895; 45; TL : Sikkim
Bertula depressalis (Snellen, [1886])
Sumatra, Bali, Peninsular Malaysia, Borneo, Philippines. See [maps ]
Bertula descripta (Leech, 1900)
Bertula erectilinea (Swinhoe, 1902)
Borneo. See [maps ]
Bertula erythrognatha Holloway, 2008 Peninsular Malaysia
Bertula erythrognatha Holloway, 2008; Moths of Borneo 17 : 35, f. 74, 134, pl. 1; TL : Penang
Bertula excelsalis Walker, 1862
Brazil. See [maps ]
Bertula excelsalis Walker, 1862; 115; TL : Brazil
Bertula figurata (Hampson, 1898)
Meghalaya. See [maps ]
Bleptina figurata Hampson, 1898; 698; TL : Khasis [Meghalaya]
Bertula fulvistrigalis Warren, 1912
Borneo. See [maps ]
Bertula fulvistrigalis Warren, 1912; 57; TL : Borneo, Mt. Marapok
Bertula gollum Holloway, 2008
Borneo. See [maps ]
Bertula gollum Holloway, 2008; Moths of Borneo 17 : 31, f. 64, pl. 1; TL : Sarawak, Gunong Mulu Nat. Park, R.G.S. Exped
Bertula grimsgaardi (Strand, 1919) Formosa
Bleptina (Bertula) grimsgaardi Strand, 1919; 161; TL : Formosa, Banshoryo distr., Sokutsu
Bertula hadenalis (Moore, 1867)
Darjeeling, Nilgiri, Formosa. See [maps ]
Herminia hadenalis Moore, 1867; 85; TL : Darjeeling
Herminia ruptistigma Hampson, 1891; 25, 100, pl. 148, f. 20; TL : Nilgiri distr.
Bleptina hadenalis var. alikangialis Strand, 1919; 160; TL : Formosa, Alikang
Bertula heteropalpia (Hampson, 1912) Madras
Bleptina heteropalpia Hampson, 1912; 1231; TL : Madras, Palnis
Bertula imparatalis Walker, [1866] Hindostan
Bertula imparatalis Walker, [1866]; 1168; TL : Hindostan
Bertula impuralis (Hampson, 1898)
Sikkim, Khasis. See [maps ]
Bleptina impuralis Hampson, 1898; 698; TL : Sikkim; Khasis
Bertula incisa (Wileman, 1915) Formosa
Bleptina incisa Wileman, 1915; 14; TL : Formosa, Arizan
Bertula inconspicua (Swinhoe, 1902)
Borneo. See [maps ]
Bertula insignifica Rothschild, 1920
Sumatra, Java, Borneo. See [maps ]
Bertula jucundalis (Walker, [1866])
Ceylon. See [maps ]
Bleptina ? jucundalis Walker, [1866]; 1164; TL : Ceylon
Bleptina jucundalis ; [POOLE]
Bertula jucundalis ; [MOB17] , 26 (implied)
Bertula kosemponica (Strand, 1917) Formosa
Bleptina kosemponica Strand, 1917; 134; TL : Formosa, Kosempo
Bertula latifasciata (Hampson, 1895)
Sikkim, Khasis, Burma. See [maps ]
Bleptina latifasciata Hampson, 1895; 43; TL : Sikkim; Khasis; Burma
Bertula lobativalva Holloway, 2008
Borneo, Singapore. See [maps ]
Bertula mesilau Holloway, 2008
Borneo. See [maps ]
Bertula mesilau Holloway, 2008; Moths of Borneo 17 : 31, f. 67, pl. 1; TL : Sabah, Mt. Kinabalu, Mesilau, 1500m
Bertula micealis (Walker, 1859)
Borneo. See [maps ]
Bleptina micealis ; [POOLE]
Bertula micealis ; [MOB17] , 37, f. 86-87, pl. 2
Bertula microdepressalis Holloway, 2008
Borneo. See [maps ]
Bertula microdepressalis Holloway, 2008; Moths of Borneo 17 : 34, f. 69, pl. 1; TL : Sarawak, Gunong Mulu Nat. Park, R.G.S. Exped.
Bertula micropulla Holloway, 2008
Borneo, Sumatra. See [maps ]
Bertula micropulla Holloway, 2008; Moths of Borneo 17 : 47, f. 97, pl. 1; TL : Sarawak, Gunong Mulu Nat. Park, R.G.S. Exped.
Bertula mimica (Hampson, 1898)
Sikkim. See [maps ]
Bleptina mimica Hampson, 1898; 698; TL : Sikkim
Bertula momusalis (Walker, [1859])
Borneo, Singapore. See [maps ]
Bertula momusalis ; [MOB17] , 28, f. 61, 123, pl. 1
Bertula nigra Swinhoe, 1902
Borneo, Singapore. See [maps ]
Bertula nigristigma (Leech, 1900)
Bleptina nigristigma ; [POOLE]
Bertula nigristigma ; [MOB17] , 26 (implied)
Bertula ningpoalis (Leech, 1889)
Herminia ningpoalis Leech, 1889; Entomologist 22 : 64 , pl. 2 , f. 7; TL : Snow Valley, near Ningpo
Bleptina ningpoalis ; [POOLE]
Bertula ningpoalis ; [MOB17] , 26 (implied)
Bertula ochracealis (Moore, 1867)
Darjeeling. See [maps ]
Herminia ochracealis Moore, 1867; 85; TL : Darjeeling
Herminia ochracealis ; [NHM card ]
Bleptina ochracealis ; [POOLE]
Bertula ochracealis ; [MOB17] , 26 (implied)
Bertula owadai Holloway, 2008
Borneo, Sumatra. See [maps ]
Bertula owadai Holloway, 2008; Moths of Borneo 17 : 39, f. 91, pl. 2; TL : Sarawak, Gunong Mulu Nat. Park, R.G.S. Exped.
Bertula pallidicosta Holloway, 2008
Borneo. See [maps ]
Bertula pallidicosta Holloway, 2008; Moths of Borneo 17 : 45, f. 98, pl. 1; TL : Sarawak, Gunong Mulu Nat. Park, R.G.S. Exped.
Bertula pallidisca Holloway, 2008
Borneo. See [maps ]
Bertula pallidisca Holloway, 2008; Moths of Borneo 17 : 32, f. 70, pl. 1; TL : Sarawak, Gunong Mulu Nat. Park, R.G.S. Exped
Bertula pallidipulla (Holloway, 1976)
Borneo. See [maps ]
Bleptina pallidipulla Holloway, 1976; Moths of Borneo with special reference to Mt. Kinabalu : 42, pl. 16, f. 190; TL : Borneo, Mt. Kinabalu, Park H.Q.
Bleptina pallidipulla ; [POOLE]
Bertula pallidipulla ; [MOB17] , 45, f. 95, pl. 1
Bertula parallela (Leech, 1900)
Bertula partita Hampson, 1891
S.India, S.China, Borneo, Bali, Sulawesi. See [maps ]
Bertula persimilis (Wileman, 1915) Formosa
Bleptina persimilis Wileman, 1915; 13; TL : Formosa, Rantaizan
Bertula phidiasalis Walker, [1859]
Borneo. See [maps ]
Bertula phidiasalis Walker, [1859]; 166; TL : Borneo, Sarawak
Bertula phlegeusalis (Walker, [1859])
Borneo. See [maps ]
Elyra phlegeusalis ; [POOLE]
Bertula phlegeusalis ; [MOB17] , 34, f. 73, 133, pl. 1
Bertula picta (Pagenstecher, 1894)
Java, Borneo. See [maps ]
Bleptina picta ; [POOLE]
Bertula picta ; [MOB17] , 43, f. 40, 68, 126, pl. 1
Bertula pollocki Holloway, 2008
Borneo. See [maps ]
Bertula pollocki Holloway, 2008; Moths of Borneo 17 : 30, f. 62, 124, pl. 1; TL : Matang, 3600ft, Borneo
Bertula postlineata (Holloway, 2008)
Borneo. See [maps ]
"Bertula" postlineata Holloway, 2008; Moths of Borneo 17 : 48, f. 102, 135, pl. 1; TL : N.Borneo, Kina Balu
Bertula proctori Holloway, 2008
Borneo. See [maps ]
Bertula proctori Holloway, 2008; Moths of Borneo 17 : 48, f. 85, pl. 1; TL : Sarawak, Gunong Mulu Nat. Park, R.G.S
Bertula propugnata (Leech, 1900)
Bleptina propugnata ; [POOLE]
Bertula propugnata ; [MOB17] , 26 (implied)
Bertula proxima (Leech, 1900)
Bertula prunosa (Moore, [1885])
Ceylon. See [maps ]
Hadennia prunosa Moore, [1885]; 237, pl. 177, f. 8; TL : Ceylon
Bertula pughi Holloway, 2008
Borneo. See [maps ]
Bertula pughi Holloway, 2008; Moths of Borneo 17 : 42, f. 80, pl. 1; TL : Sarawak, Gunong Mulu Nat. Park, R.G.S. Exped.
Bertula pulla (Swinhoe, 1902)
Borneo, .... See [maps ]
Bertula quadripuncta (Wileman, 1915) Formosa, ...
Bleptina quadripuncta Wileman, 1915; 15; TL : Formosa, Kanshirei
Bleptina quadripuncta ; [POOLE]
Bertula quadripuncta ; [MOB17] , 26 (implied)
Bertula rectilinea (Leech, 1900)
Bleptina rectilinea ; [POOLE]
Bertula rectilinea ; [MOB17] , 26 (implied)
Bertula restricta (Moore, 1882)
Darjeeling. See [maps ]
Herminia restricta Moore, 1882; 192; TL : Darjeeling
Bertula rostrilinea Prout, 1928
Borneo. See [maps ]
Bertula rufithorax Holloway, 2008
Borneo. See [maps ]
Bertula rufithorax Holloway, 2008; Moths of Borneo 17 : 32, f. 66, pl. 1; TL : Sabah, Mt. Kinabalu, Park H.Q., 1620m
Bertula rufosa (Holloway, 1976)
Borneo. See [maps ]
Badiza rufosa Holloway, 1976; 42, pl. 15, f. 188; TL : Borneo, Mt. Kinabalu, Park H.Q.
Bertula saigonensis Lemee, 1950
Vietnam. See [maps ]
Bertula saigonensis Lemee, 1950; 72; TL : Saigon
Bertula scoonesi Holloway, 2008
Borneo. See [maps ]
Bertula scoonesi Holloway, 2008; Moths of Borneo 17 : 41, f. 81, pl. 1; TL : Sarawak, Gunong Mulu Nat. Park, R.G.S. Exped.
Bertula scoulari Holloway, 2008
Borneo. See [maps ]
Bertula scoulari Holloway, 2008; Moths of Borneo 17 : 41, f. 79, pl. 1; TL : Sarawak, Gunong Mulu Nat. Park, R.G.S. Exped.
Bertula serratula (Bethune-Baker, 1908)
New Guinea. See [maps ]
Bleptina serratula Bethune-Baker, 1908; 207; TL : British New Guinea, Mt Kebea
Bertula sinuosa (Leech, 1900)
Japan, China. See [maps ]
Bertula spacoalis (Walker, 1859)
Japan, Korea, NE.China. See [maps ]
Bertula sphaerula (Swinhoe, 1902)
Borneo. See [maps ]
Bocana sphaerula ; [POOLE]
Bertula sphaerula ; [MOB17] , 42, f. 84, pl. 1
Bertula spineusalis Holloway, 2008
Borneo. See [maps ]
Bertula spineusalis Holloway, 2008; Moths of Borneo 17 : 35, f. 77, pl. 1; TL : Sarawak, Gunong Mulu Nat. Park, R.G.S. Exped.
Bertula subnigripuncta Holloway, 2008
Borneo. See [maps ]
Bertula subnigripuncta Holloway, 2008; Moths of Borneo 17 : 33, f. 71, pl. 1; TL : Sarawak, Gunong Mulu Nat. Park, R.G.S. Exped.
Bertula suisharyonis (Strand, 1920) Formosa
Hypena ? suisharyonis Strand, 1920; 172; TL : Formosa, Suisharyo
Bertula terminalis (Wileman, 1915) Formosa
Bleptina terminalis Wileman, 1915; 15; TL : Formosa, Arizan
Bertula tespisalis (Walker, [1859])
Borneo, Peninsular Malaysia. See [maps ]
Bertula transcaerulea Holloway, 2008
Borneo. See [maps ]
Bertula transcaerulea Holloway, 2008; Moths of Borneo 17 : 38, f. 89, pl. 2; TL : Bidi, Sarawak
Bertula tripartita (Leech, 1900)
Bleptina tripartita ; [POOLE]
Bertula tripartita ; [MOB17] , 26 (implied)
Bertula vialis (Moore, 1882)
Meghalaya. See [maps ]
Herminia vialis Moore, 1882; 192; TL : Cherra Punji [Meghalaya]
Bertula venata (Leech, 1900)
Sichuan, ... See [maps ]
Bertula vestitalis (Leech, 1900)
W.China. See [maps ]
Bleptina vestitalis ; [POOLE]
Bertula vestitalis ; [MOB17] , 26 (implied)
Bertula wallacea Holloway, 2008
Boreno, Sulawesi, Seram, Ambon. See [maps ]
Bertula wallacea Holloway, 2008; Moths of Borneo 17 : 36, f. 45, 76, 132, pl. 1; TL : Brunei, 1618m, Bukit Retak
Bertula waterstradti Holloway, 2008
Borneo, NE.Himalaya (Sikkim). See [maps ]
Bertula waterstradti Holloway, 2008; Moths of Borneo 17 : 37, f. 125, pl. 1; TL : N.Borneo, Mt. Kina Balu
Bertula willotti Holloway, 2008
Borneo. See [maps ]
Bertula willotti Holloway, 2008; Moths of Borneo 17 : 36, f. 75, pl. 1; TL : Malaysia, Sabah, Danum Valley, 170m, 4°58'N 117°48'E
Unknown or unplaced taxa
Bleptina histaspesalis ; [POOLE]
"Bleptina" histaspesalis ; [MOB17] , 215, pl. 9
3.7.2019 (96)
Some related literature:
Felder & Rogenhofer, 1874
Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Behilfen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil. Band 2. Abtheilung 2. Lepidoptera. Rhopalocera
Reise Fregatte Novara, Bd 2 (Abth. 2)
(1): 1 -136, pl. 1 -21 (Rhop.) (C. & R. Felder, [1865]),
(2): 137 -378, pl. 22 -47 (Rhop.) (C. & R. Felder, 1865),
(3): 379 -536, pl. 48 -74 (Rhop.) (C. & R. Felder, 1867),
(4): 537 -548, pl. 75 -120, (Inhalts-Verz.) 1-9 (pl. 1-74), (Felder & Rogenhofer, 1874) ,
(5): pl. 121 -140, (erklärung) 1 -10 (pl. 75-107), (Inhalts-Verz.) 1-20 (pl. 108-140), (Felder & Rogenhofer, 1875)
Hampson, 1891
Illustrations of typical specimens of Lepidoptera Heterocera in the collection of the British Museum. Part 8. The lepidoptera of heterocera of the Nilgiri district
Ill. typical Spec. Lep. Het. Colln Br. Mus. 8
: 1 -144, pl. 139 -156
Hampson, 1925
Two new genera and a new species of oriental Hypeninæ (Noctuidæ)
Ann. Mag. nat. Hist. (9) 15 (87)
: 408-411
[MOB17]; Holloway, 2008
Family Noctuidae, subfamilies Rivulinae, Phytometrinae, Herminiinae, Hypeninae and Hypenodinae
Moths of Borneo 17
: 1-268
Leech, 1889
New species of Deltoids and Pyrales from Corea, North China, and Japan
Entomologist 22
: 62 -71, pl. 2 -4
Leech, 1900
Lepidoptera Heterocera from Northern China, Japan, and Corea. Part III, IV
Trans. Ent. Soc. Lond. 1900
(1): 9 -161,
(4): 511 -663
[RFEL]; Leley, 2016
Annotated catalogue of the insects of Russian Far East. Volume II. Lepidoptera
Cat. ins. Russian Far East. 2
: 1-812
Owada, 1987
A taxonomic study on the subfamily Herminiinae of Japan : (Lepidoptera, Noctuidae)
Nat. Sci. Mus. monogr. 2 : 1-208
Pagenstecher, 1894
Beiträge zur Lepidopteren - Fauna des Malayischen Archipels. (9 & 10)
Jb. Nassau. Ver. Nat. 47
: 25 -58, pl. 1,
: 59 -81, pl. 2-3
[POOLE]; Poole, 1989
Lepidopterorum catalogus (New Series) 118: Noctuidae
Lepid. cat. (n.s.) 118 :
Prout, 1928
Noctuid moths from some of the mountains of Sarawak
Sarawak Mus. J. 3
: 461 -503, pl. 15 -16
Rothschild, 1920
Heterocera. Collected in Korinchi, West Sumatra, by Messrs. H.C. Robinson and C. Boden Kloss
J. Fed. Malay States Mus. 8 (3)
: 107-139
Snellen, [1886]
Natuurlijke Historie. Achtse afdeeling. Lepidoptera door P. C. T. Snellen, met eene inleiding door Joh. F. Snelleman. Midden-Sumatra
Midden-Sumatra, 4 (8)
: 1 -84, pl. 1 -2 (?1880, ?1884, [1886]),
: 85 -92, pl. 3 -5 ([1892])
Swinhoe, 1902
Descriptions of new Eastern and Australian Moths
Ann. Mag. nat. Hist. (7) 9 (51)
: 165 -182
Swinhoe, 1902
New Eastern Heterocera
Ann. Mag. nat. Hist. (7) 10 (60)
: 495 -503
Swinhoe, 1906
Eastern and African Heterocera
Ann. Mag. Nat. Hist. (7) 17 (102)
: 540 -556
Walker, 1859
List of the Specimens of Lepidopterous Insects in the Collection of the British Museum. Supplement
List Spec. Lepid. Insects Colln Br. Mus.
16 : 1 -253 ([1859]),
17 : 255 -508 (1859),
18 : 509 -798 (1859),
19 : 799 -1036 (1859)
Walker, [1863]
Catalogue of the Heterocerous Lepidopterous Insects collected at Sarawak, in Borneo, by Mr. A. R. Wallace, with Descriptions of New Species
J. Proc. Linn. Soc. (Zool.) 7
: 49 -84, 160 -198
If you have corrections, comments or
information to add into these pages, just send mail to
Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.