Erynnis

Erynnis Schrank, 1801 Duskywings

Erynnis Schrank, 1801; Fauna Boica 2 (1): 152, 157; TS: Papilio tages Linnaeus
Thymele Fabricius, 1807; Mag. f. Insektenk. 6: 287; TS: Papilio tages Linnaeus
Thymale Oken, 1815; Lehrbuch Naturgesch. 3 (Zool.) (1): 757-758 (unavail.); TS: Papilio tages Linnaeus
Astycus Hübner, 1822; Syst.-alph. Verz.: 1, 3, 5, 6, 8-10; TS: Papilio tages Linnaeus
Thanaos Boisduval, [1834]; Icon. hist. Lépid. Europe 1 (23-24): 240; TS: Papilio tages Linnaeus
Thanatos Dunning & Pickard, 1858; Accent. List Brit. Lep.: 6 (unjust. emend.)
Hallia Tutt, 1906; Nat. Hist. Br. Lepid. 8: 261 (preocc. Hallia Edwards & Haime, 1850); TS: Thanaos marloyi Boisduval
Erynnides Burns, 1964

Thanaos; Godman & Salvin, [1899], Biol. centr.-amer., Lep. Rhop. 2: 455
Erynnis (Pyrgini); [NL4A], 47
Erynnis (Pyrginae); Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 8; [RFEL]

900x617 (~75Kb) USA: Near Crystal Mtn, (46°57'51N 121°28'55W), Pierce Co., WA, 27.7.1999, Photo © Markku Savela

700x652 (~66Kb) USA: Near Crystal Mtn, (46°57'51N 121°28'55W), Pierce Co., WA, 27.7.1999, Photo © Markku Savela

identification uncertain 500x378 (~51Kb) Venezuela, Parai Tepui near Mount Roraima in the Gran Sabana, July 2002, Photo © Simon Coombes

The exact identification of these species is still unknown, but tentatively assumed to belong into this group.

[NL4A] introduced subgeneric division, but I don't know how to place the species outside neotropics.

Erynnis montanus (Bremer, 1861) S.China - Amur, Japan. See [maps]
- Hesperia montanus Bremer, 1861; Bull. Acad. Imp. Sci. St. Petersb. 3 Mélanges biol. St.-Pétersb. 3 (5-6): 556; TL: Bureinsky Mts., Amur region
- Thanaos rusticanus Butler, 1866; J. Linn. Soc. Zool. Lond. 9 (34): 58; TL: Japan, Hakodate
- Erynnis montanus ; [BOW]: pl. 208, f. 9
- Erynnis montanus, Lepidoptera collection [SZMN]
- Erynnis mountanus[sic, recte montanus]; Kudrna, 1974, Atalanta 5: 116
- Erynnis montanus; [BRU], 106; [MRS], 705; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 8; [RFEL]
Larva on Quercus mongolica [BRU]
- E. m. nigrescens (Leech)
  - Erynnis montanus nigrescens; [MRS], 705
- E. m. monta Evans
  - Erynnis montanus monta; [MRS], 705
- E. m. neomontanus Murayama & Yoshisaka, 1959 Formosa
  - Erynnis montanus neomontanus Murayama & Yoshisaka, 1959; Tyô to Ga 10 (1): 14; TL: Poli?, Formosa

Erynnis popoviana (Nordmann, 1851)

Mongolia, N.China, E.China, C.China, Ussuri. See [maps]

Hesperia popoviana Nordmann, 1851; Bull. Soc. imp. Nat. Moscou 24: 443, pl. 12, f. 3-4; TL: Kyakhta, Transbaikalia
Nisoniades tages var. sinina Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4): 461

Erynnis tages popoviana; [MRS], 705
Erynnis popoviana; [BRU], 107; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 8; [RFEL]

800x600 (~121Kb) upperside Wet ground at the pool in the pine forest called Tsasucheiskii Bor at the village Nizhnii Tsasuchei, Onon District, Chita Province, SE Transbaikalia, Siberia, Russia. 22nd June 1995, Photo © Oleg Kosterin

Erynnis pathan Evans, 1949 Ghissar - Chitral, N.Baluchistan. See [maps]
- Erynnis pathan Evans, 1949; Cat. Hesp. Europe Asia Australia Brit. Mus.: 168; TL: Chitral, Utzun Valley
- Erynnis pathan; [BRU], 107; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 8
Erynnis pelias (Leech, 1891) W.China. See [maps]
- Nisoniades pelias Leech, 1891; Entomologist 24 (Suppl.): 60; TL: Ta-chein-lu; Wa-ssu-kou
- Nisoniades erebus Grum-Grshimailo, 1891; Horae Soc. ent. Ross. 25 (3-4): 461
- Erynnis pelias erebus; [NHM card]
- Erynnis pelias; Huang, 2001

Subgenus Erynnis Schrank, 1801

Erynnis Schrank, 1801; Fauna Boica 2 (1): 152, 157; TS: Papilio tages Linnaeus
Thymele Fabricius, 1807; Mag. f. Insektenk. 6: 287; TS: Papilio tages Linnaeus
Thymale Oken, 1815; Lehrbuch Naturgesch. 3 (Zool.) (1): 757-758 (unavail.); TS: Papilio tages Linnaeus
Astycus Hübner, 1822; Syst.-alph. Verz.: 1, 3, 5, 6, 8-10; TS: Papilio tages Linnaeus
Thanaos Boisduval, [1834]; Icon. hist. Lépid. Europe 1 (23-24): 240; TS: Papilio tages Linnaeus
Thanatos Dunning & Pickard, 1858; Accent. List Brit. Lep.: 6 (unjust. emend.)
Hallia Tutt, 1906; Nat. Hist. Br. Lepid. 8: 261 (preocc. Hallia Edwards & Haime, 1850); TS: Thanaos marloyi Boisduval

Erymnis; Rafinesque, 1815, (missp.)
Erinnys; Ochsenheimer, 1816, (missp.)
Tanaus; Blanchard, 1840, (missp.)
Thannaos; Lang, 1884, (missp.)
Thanaus; Kirby, 1897, in Allen, Nat. Libr., Lepid. 3: 12 (missp.)
Thymela; Raymundo da Silva, 1907, (missp.)
Thanaes; Simms, 1920, (missp.)
Thimele; Hayward, 1933, (missp.)
Astyeus; Evans, 1953, (missp.)
Thymate; Hayward, 1973, (missp.)
Prynnis; Miller, 1992, (missp.)
Thymeles; Nekrutenko, 2003, (missp.)
Erynnis (Erynnis); [NL4A], 47

Erynnis tages (Linnaeus, 1758)

CEU, SEU, tr, Russia, Siberia, C.Asia, China, Amur. See [maps] Dingy Skipper Dunkler Dickopffalter Skogssmygare La Grisette Cervantes Powszelatek brunatek

Papilio tages Linnaeus, 1758; Syst. Nat. (Edn 10) 1: 485
morio (Scopoli, 1763)
geryon (Rottenburg, [1775])

Erynnis tages, Lepiforum [LEPIFORUM]
Erynnis tages, Schmetterlinge der Schweiz [Heiner Ziegler]
Erynnis tages, Moths and Butterflies of Europe [MBE]
Erynnis tages, Larvae of North-European Lepidoptera [Kimmo Silvonen]
Erynnis tages, Svenska fjärilar [SVF]
Erynnis tages, Captain's European Butterfly Guide [Simon Coombes]

Erynnis tages; [BOW]: pl. 11, f. 15; [BRU], 106; [MRS], 705; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 8; Yakovlev, 2012, Nota lepid. 35 (1): 60; [RFEL]

500x319 (~36Kb) upperside Haldon woods, Devon, UK 21/05/95, Photo © Simon Coombes

800x600 (~114Kb) upperside Open pine forest on a hill just NE of the Chita City, Transbaikalia, Siberia, Russia. 17th June 1995, Photo © Oleg Kosterin

Larva on Lotus corniculatus, Eryngium , Coronilla [H&R], Medicago , Hippocrepis [BRU]

E. t. unicolor Freyer, 1852 Transcaucasus
E. t. elbursina (Bytinski-Salz & Brandt, 1937) Iran
- Thanaos elbursina Bytinski-Salz & Brandt, 1937; Ent. Rec. J. Var. (Suppl.) 49: 3

Erynnis marloyi (Boisduval, [1834])

Balkans, Asia Minor, Syria, Iran, Chitral. See [maps] Inky Skipper

Thanaos marloyi Boisduval, [1834]; Icon. hist. Lépid. Europe 1 (23-24): 241, pl. 47, f. 6-7; TL: Peloponnesos, Greece
sericea (Freyer, [1838])
rustan (Kollar, [1849])

Erynnis marloyi, Lepiforum [LEPIFORUM]
Erynnis marloyi, Moths and Butterflies of Europe [MBE]

Erynnis marloyi; [BOW]: pl. 11, f. 14; [BRU], 107; [Otakar Kudrna]; Korb & Bolshakov, 2011, Eversmannia Suppl. 2: 8

1000x1212 (~415Kb) Iran, Ostan-e Markazi, Shakhrestan-e Shazand, Astaneh Town env., Seidan chashmah, 2289 m a.s.l., 20th May 2017, Photo © Oleg Kosterin

1000x1121 (~410Kb) Iran, Ostan-e Markazi, Shakhrestan-e Shazand, Astaneh Town env., Seidan chashmah, 2289 m a.s.l., 20th May 2017, Photo © Oleg Kosterin

Erynnis brizo (Boisduval & Le Conte, [1834]) E.USA, Colorado, Arizona. See [maps] Sleepy Duskywing
- Thanaos brizo Boisduval & Le Conte, [1834]; Hist. Lép. Am. Sept. (23-26): pl. 66; TL: N.Amer.
- Erynnis brizo, Moth Photographers Group [MPG]
- Erynnis brizo brizo, Butterflies of America [BOA]
- Erynnis brizo, Butterflies and Moths of North America [BAMONA]
- Thanaos brizo; Dyar, 1903, Bull. U.S. nat. Mus. 52: 59
- Erynnis brizo; [NACL], #3946; [Opler]
- Erynnis (Erynnis) brizo; [NL4A], #751
Larva on Quercus , less Castanea dentata [NAB]
- E. b. brizo Mexico, ...
  - Erynnis briso; Okano, 1981, (missp.)
  - Erynnis (Erynnis) brizo brizo; [NL4A], #751a
- E. b. somnus (Lintner, 1881) Florida?
  - Nisoniades[?] somnus Lintner, 1881; Papilio 1 (5): 73
  - Erynnis brizo somnus, Butterflies of America [BOA]
  - Thanaos somnus; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
  - Erynnis brizo somnus; [NACL], #3946a
- E. b. lacustra (Wright, 1905) Mexico, California
  - brizo lacustra (Wright, 1905) (Nisoniades); ; TL: California
  - Erynnis brizo lacustra, Butterflies of America [BOA]
  - Erynnis pacuvius lacustra Wright, 1904;
  - Erynnis brizo lacustra; [NACL], #3946c
  - Erynnis (Erynnis) brizo lacustra; [NL4A], #751c
- E. b. burgessi (Skinner, 1914) Mexico, Arizona
  - brizo burgessi (Skinner, 1914) (Thanaos); ; TL: Arizona
  - Erynnis brizo burgessi, Butterflies of America [BOA]
  - Erynnis brizo burgessi, Butterflies of Southeastern Arizon [Bruce Walsh]
  - Erynnis burgessii; Klots, 1951, (missp.)
  - Erynnis brizo burgessi; [NACL], #3946b
  - Erynnis (Erynnis) brizo burgessi; [NL4A], #751b
- E. b. muelleri (Draudt, 1923) Mexico
  - brizo muelleri (Draudt, 1923) (Thanaos); ; TL: Mexico, Puebla
  - Erynnis brizo muelleri, Butterflies of America [BOA]
  - Thanaos mulleri; Hoffmann, 1932, (missp.)
  - Erynnis (Erynnis) brizo muelleri; [NL4A], #751d
Erynnis icelus (Scudder & Burgess, 1870) New York, ..?.., California. See [maps] Dreamy Duskywing Aspen Dusky Wing
- Nisoniades icelus Scudder & Burgess, 1870; ; TL: "New England"
- Nisoniades icelus Lintner, 1872; 23 Rept. N.Y. Sta. Cab. Nat. Hist.: 162 (preocc.); TL: New York
- Nisoniades bautista Plötz, 1884; ; TL: California
  - = ; [NL4A], #752
- Nisoniades hamamaelidis Scudder, 1889; (nom. nud.); TL: USA
  - = ; [NL4A], #752
- Erynnis icelus, Moth Photographers Group [MPG]
- Erynnis icelus, Butterflies of America [BOA]
- Erynnis icelus, Butterflies and Moths of North America [BAMONA]
- Erynnis icelus, Butterflies of Southeastern Arizon [Bruce Walsh]
- ?Thanaos icelus Lintner, 1872; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- Erynnis hamamaelides; Bridges, 1983, (missp.)
- Erynnis icelus; [NACL], #3945; [BOW]: pl. 21, f. 36; [BOC], 51; [Opler]
- Erynnis (Erynnis) icelus; [NL4A], #752
Larva on Salix , Populus , Robinia , Betula [NAB]
Erynnis mercurius (Dyar, 1926) Mexico. See [maps]
- Thanaos mercurius Dyar, 1926; ; TL: Mexico, Distrito Federal
- Erynnis mercurius, Butterflies of America [BOA]
- Erynnis (Erynnis) mercurius; [NL4A], #753

Subgenus Erynnides Burns, 1964

Erynnides (Erynnis); [NL4A], 47

Erynnis afranius (Lintner, 1878) Mexico - Manitoba. See [maps] Afranius Duskywing Bald Duskywing
- Nisoniades afranius Lintner, 1878; 30 Rept. N.Y. Mus. Nat. Hist.: 175; TL: "Colorado"
- Erynnis afranius, Moth Photographers Group [MPG]
- Erynnis afranius, Butterflies of America [BOA]
- Erynnis afranius, Butterflies and Moths of North America [BAMONA]
- Erynnis afranius, Butterflies of Southeastern Arizon [Bruce Walsh]
- Thanaos afranius; Godman & Salvin, [1899], Biol. centr.-amer., Lep. Rhop. 2: 459, 3 pl. 91, f. 24-26; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- Erynnis afranius; [NACL], #3960; [BOC], 54; [Opler]
- Erynnis (Erynnides) afranius; [NL4A], #754
Larva on Lupinus , Lotus [NAB], Thermopsis rhombifolia [BOC]
Erynnis funeralis (Scudder & Burgess, 1870) California, S.Nevada, S.Utah, Colorado, W.Kansas - Texas - Argentina, Chile, Colombia. See [maps] Funereal Duskywing
- Nisoniades funeralis Scudder & Burgess, 1870; Proc. Bost. Soc. Nat. Hist. 13: 293; TL: Texas
- Helias clericalis Burmeister, 1875; ; TL: Argentina
  - = ; [NL4A], #755
- Nisoniades australis Mabille, 1883; Bull. Soc. Ent. Belg. 27: liv [54]; TL: Colombia
  - = ; [NL4A], #755
- Erynnis funeralis, Moth Photographers Group [MPG]
- Erynnis funeralis, Butterflies of America [BOA]
- Erynnis funeralis, Butterflies and Moths of North America [BAMONA]
- Erynnis funeralis, Butterflies of Southeastern Arizon [Bruce Walsh]
- Phlebodes clericalis; Burmeister, 1878
- Erynnis zarucco funeralis;
- Thanaos funeralis; Godman & Salvin, [1899], Biol. centr.-amer., Lep. Rhop. 2: 456, 3 pl. 91, f. 10-14; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- Erynnis funeralis; [NACL], #3957; [BOW]: pl. 21, f. 34; [Opler]; Benyamini, Ugarte, Shapiro, Mielke, Pyrcz & Bálint, 2014, Bol. Mus. Nac. Hist. Nat. Chile 63: 21 (list)
- Erynnis (Erynnides) funeralis; [NL4A], #755
Larva on Fabaceae [NAB]
Erynnis horatius (Scudder & Burgess, 1870) E.USA, Florida, Mexico. See [maps] Horace's Duskywing
- Nisoniades horatius Scudder & Burgess, 1870; Proc. Bost. Soc. Nat. Hist. 13: 301; TL: Massachusetts/Texas
- Nisoniades virgilius Scudder & Burgess, 1870; ; TL: USA
  - = ; [NL4A], #756
- Nisoniades petronius Mead, 1875; (nom. nud.)
- Nisoniades petronius Lintner, 1881; Papilio 1 (5): 70; TL: Florida
  - = ; [NL4A], #756
- Erynnis horatius, Moth Photographers Group [MPG]
- Erynnis horatius, Butterflies of America [BOA]
- Erynnis horatius, Butterflies and Moths of North America [BAMONA]
- Thanaos petronius; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- Thanaos horatius; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- Thanaos horatuis; Draudt, 1923, (missp.)
- Erynnis horatius; [NACL], #3952; [BOW]: pl. 21, f. 35; [Opler]
- Erynnis (Erynnides) horatius; [NL4A], #756
Larva on Quercus muehlenbergii, Q. gambelii, Q. texana [NAB]
Erynnis juvenalis (Fabricius, 1793) E.USA - Rocky Mts.. See [maps] Juvenal's Duskywing
- Hesperia juvenalis Fabricius, 1793; Ent. Syst. 3: 339; TL: America
- Erynnis juvenalis, Moth Photographers Group [MPG]
- Erynnis juvenalis juvenalis, Butterflies of America [BOA]
- Erynnis juvenalis, Butterflies and Moths of North America [BAMONA]
- Thanaos juvenalis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- Erynnis juvenalis; [NACL], #3947; [BOW]: pl. 21, f. 35 (text); [Opler]
- Erynnis (Erynnides) juvenalis; [NL4A], #757
Larva on Quercus velutina, Q. rubra, Q. alba [NAB]
- E. j. juvenalis Florida, ...?
  - Nisoniades juvenis Hübner, [1819]; (repl. name); TL: America
    
    = Thanaos juvenalis ; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
    = Erynnis juvenalis juvenalis ; [NL4A], #757a
  - Nisoniades costalis Westwood, 1852
    
    = Erynnis juvenalis juvenalis ; [NL4A], #757a
  - Nisoniades ennius Scudder & Burgess, 1870; ; TL: USA
    
    = Erynnis juvenalis juvenalis ; [NL4A], #757a
  - Nisoniades plautus Scudder & Burgess, 1870; Proc. Bost. Soc. Nat. Hist. 13: 304; TL: Florida
    
    = Erynnis juvenalis juvenalis ; [NL4A], #757a
  - Erynnis juvenalis plautus;
  - Thanaos plautus; Dyar, 1903, Bull. U.S. nat. Mus. 52: 61
  - Erynnis (Erynnides) juvenalis juvenalis; [NL4A], #757a
- E. j. clitus (Edwards, 1883) Arizona, New Mexico, Mexico
  - Thanaos clitus Edwards, 1882; Papilio 2 (9-10): 180; TL: Arizona
  - Thanaos maestus Godman, [1899]; Biol. centr.-amer., Lep. Rhop. 2: 457, 3 pl. 91, f. 18; TL: Arizona; Mexico, Pinal, Puebla
    
    = Erynnis juvenalis clitus ; [NL4A], #757b
  - Erynnis juvenalis clitus, Butterflies of America [BOA]
  - Erynnis juvenalis clitus, Butterflies of Southeastern Arizon [Bruce Walsh]
  - Thanaos clitus; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
  - Erynnis clitus; [BOW]: pl. 21, f. 34 (text)
  - Nisoniades moestus; Skinner, 1904, (missp.)
  - Erynnis juvenalis clitus; [NACL], #3947a
  - Erynnis (Erynnides) juvenalis clitus; [NL4A], #757b
Erynnis martialis (Scudder, 1869) NE.USA - Alabama. See [maps] Mottled Duskywing
- Nisoniades martialis Scudder, 1869; Trans. Chicago Acad. Sci. 1 (2): 355; TL: Iowa
- Thanaos quercus Butler, 1870; (nom. nud.)
  - = ; [NL4A], #758
- Nisoniades ausonius Lintner, 1872; 23 Rept. N.Y. Sta. Cab. Nat. Hist.: 166; TL: New York
  - = ; [NL4A], #758
- Erynnis martialis, Moth Photographers Group [MPG]
- Erynnis martialis, Butterflies of America [BOA]
- Erynnis martialis, Butterflies and Moths of North America [BAMONA]
- Thanaos ausonius; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- Thanaos martialis; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- Erynnis martialis; [NACL], #3954; [BOW]: pl. 21, f. 37; [Opler]
- Erynnis (Erynnides) martialis; [NL4A], #758
Larva on Ceanothus americanus, C. fendleri [NAB]
Erynnis meridianus Bell, 1927 California, Nevada, New Mexico, .... See [maps] Meridian Duskywing Southern Duskywing
- Erynnis meridianus Bell, 1927; J. N.Y. Ent. Soc. 35 (3): 261; TL: Arizona
- Erynnis meridianus, Moth Photographers Group [MPG]
- Erynnis meridianus meridianus, Butterflies of America [BOA]
- Erynnis meridianus, Butterflies and Moths of North America [BAMONA]
- Erynnis meridianus, Butterflies of Southeastern Arizon [Bruce Walsh]
- Erynnis meridianus; [NACL], #3950; [BOW]: pl. 21, f. 35 (text); [Opler]
- Erynnis (Erynnides) meridianus; [NL4A], #759
- E. m. meridianus Arizona, Mexico, ...
  - Erynnis meridanus; Ferris, 1976, (missp.)
  - Erynnis (Erynnides) meridianus meridianus; [NL4A], #759a
- E. m. fieldi Burns, 1964 Mexico
  - Erynnis meridianus fieldi Burns, 1964; ; TL: Mexico, Michoacán
  - Erynnis meridianus fieldi, Butterflies of America [BOA]
  - Erynnis (Erynnides) meridianus fieldi; [NL4A], #759b
Erynnis pacuvius (Lintner, 1878) W.USA, S.USA. See [maps] Pacuvius Duskywing Dyar's Duskywing Buckthorn Dusky Wing
- Nisoniades pacuvius Lintner, 1878; 30 Rept. N.Y. Mus. Nat. Hist.: 172; TL: New Mexico
- Erynnis pacuvius, Moth Photographers Group [MPG]
- Erynnis pacuvius pacuvius, Butterflies of America [BOA]
- Erynnis pacuvius, Butterflies and Moths of North America [BAMONA]
- Erynnis pacuvius pacuvius, Butterflies of Southeastern Arizon [Bruce Walsh]
- Thanaos pacuvius; Godman & Salvin, [1899], Biol. centr.-amer., Lep. Rhop. 2: 458, 3 pl. 91, f. 16-17; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- Erynnis pacuvius; [NACL], #3955; [BOW]: pl. 21, f. 33 (text); [BOC], 53; [Opler]
- Erynnis (Erynnides) pacuvius; [NL4A], #760
Larva on Ceanothus , C. fendleri [NAB]
- E. p. pacuvius Mexico, New Mexico
  - Erynnis (Erynnides) pacuvius pacuvius; [NL4A], #760a
- E. p. lilius (Dyar, 1904) British Columbia, Washington
  - pacuvius lilius (Dyar, 1904); ; TL: Kaslo, British Columbia
  - Erynnis pacuvius lilius, Butterflies of America [BOA]
  - Erynnis pacuvius lilius; [NACL], #3955a; [BOC], 53
  Larva on Ceanothus velutinus [BOC]
- E. p. pernigra (Grinnell, 1905)
  - Erynnis pacuvius pernigra, Butterflies of America [BOA]
  - ?Erynnis persius pernigra;
  - Erynnis pacuvius perniger[sic]; [NACL], #3955b
- E. p. callidus (Grinnell, 1904) California, Mexico
  - Thanaus callidus Grinnell, 1905; ; TL: California
  - Erynnis pacuvius callidus, Butterflies of America [BOA]
  - Erynnis pacuvius callidus; [NACL], #3955c
  - Erynnis (Erynnides) pacuvius callidus; [NL4A], #760b
Erynnis propertius (Scudder & Burgess, 1870) from (British Columbia - Baja California) - Nevada. See [maps] Propertius Duskywing Western Oak Dusky Wing
- Nisoniades propertius Scudder & Burgess, 1870; Proc. Bost. Soc. Nat. Hist. 13: 298; TL: California
- Nisoniades tibullus Scudder & Burgess, 1870; Proc. Bost. Soc. Nat. Hist. 13: 298; TL: California
  - = ; [NL4A], #761
- ?Nisoniades[?] propertius Lintner, 1881; Papilio 1 (5): 71
- Erynnis propertius, Moth Photographers Group [MPG]
- Erynnis propertius, Butterflies of America [BOA]
- Erynnis propertius, Butterflies and Moths of North America [BAMONA]
- Thanaos propertius Lintner, 1881; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- Thanaos tibullus; Dyar, 1903, Bull. U.S. nat. Mus. 52: 61
- Erynnis propertius; [NACL], #3949; [BOW]: pl. 21, f. 35 (text); [Opler]
- Erynnis (Erynnides) propertius; [NL4A], #761
Larva on Quercus agrifolia, Q. garryana [NAB]
Erynnis scudderi (Skinner, 1914) Mexico, Arizona. See [maps] Scudder's Duskywing
- Thanaos scudderi Skinner, 1914; ; TL: Arizona
- Erynnis scudderi, Moth Photographers Group [MPG]
- Erynnis scudderi, Butterflies of America [BOA]
- Erynnis scudderi, Butterflies and Moths of North America [BAMONA]
- Erynnis scudderi, Butterflies of Southeastern Arizon [Bruce Walsh]
- Erynnis scudderi; [NACL], #3951; [Opler]
- Erynnis (Erynnides) scudderi; [NL4A], #762
Erynnis telemachus Burns, 1960 Arizona. See [maps] Rocky Mountain Duskywing
- Erynnis telemachus Burns, 1960; Wasmann J. Biol. 18 (1): 148; TL: Arizona
- Erynnis telemachus, Moth Photographers Group [MPG]
- Erynnis telemachus, Butterflies of America [BOA]
- Erynnis telemachus, Butterflies and Moths of North America [BAMONA]
- Erynnis telemachus; [NACL], #3948; [Opler]
- Erynnis (Erynnides) telemachus; [NL4A], #763
Larva on Quercus gambelii [NAB]
Erynnis tristis (Boisduval, 1852) Arizona, New Mexico, Texas, Mexico - Colombia, CA. See [maps] Mournfull Duskywing
- Thanaos tristis Boisduval, 1852; Ann. Soc. ent. Fr. (2) 10 (2): 311; TL: California
- Erynnis tristis, Moth Photographers Group [MPG]
- Erynnis tristis tristis, Butterflies of America [BOA]
- Erynnis tristis, Butterflies and Moths of North America [BAMONA]
- Nisoniades tristis; Scudder & Burgess, 1870, Proc. Bost. Soc. Nat. Hist. 13: 303
- Thanaos tristis; Godman & Salvin, [1899], Biol. centr.-amer., Lep. Rhop. 2: 457, 3 pl. 91, f. 15; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- Erynnis tristis; [NACL], #3953; [Opler]
- Erynnis (Erynnides) tristis; [NL4A], #764
Larva on Quercus [NAB]
- E. t. tristis Mexico, California, ...
  - Erynnis tristus; Ferris, 1977, (missp.)
  - Erynnis tritus; Ferris, 1977, (missp.)
  - Erynnis ristis; Warren, 2000, (missp.)
  - Erynnis (Erynnides) tristis tristis; [NL4A], #764a
- E. t. tatius (Edwards, 1882) Arizona, Mexico, Guatemala, Costa Rica, Panama - Colombia
  - Thanaos tatius Edwards, 1882; Papilio 2 (9-10): 179; TL: [Arizona]
  - Thanaos albomarginatus Godman, [1899]; Biol. centr.-amer., Lep. Rhop. 2: 458, 3 pl. 91, f. 19-23; TL: Mexico, Cordova; Guatemala, Chisoy Valley; Costa Rica, Irazu; Panama, Chiriqui; Colombia
    
    = Erynnis tristis tatius ; [NL4A], #764c
  - Erynnis tristis tatius, Butterflies of America [BOA]
  - Erynnis tristis tatius, Butterflies of Southeastern Arizon [Bruce Walsh]
  - Nisoniades tacitus; Maynard, 1901, (missp.)
  - Thanaos tatius; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
  - Erynnis albomarginatus; [BOW]: pl. 21, f. 33
  - Erynnis tatitus; Ferris, 1977, (missp.)
  - Erynnis tristis tatius; [NACL], #3953a
  - Erynnis (Erynnides) tristis tatius; [NL4A], #764c
- E. t. pattersoni Burns, 1964 Mexico(Baja California Sur)
  - Erynnis tristis pattersoni Burns, 1964; ; TL: Mexico(Baja California Sur)
  - Erynnis tristis pattersoni, Butterflies of America [BOA]
  - Erynnis (Erynnides) tristis pattersoni; [NL4A], #764b
Erynnis zarucco (Lucas, 1857) Cuba, Texas, Florida. See [maps] Zarucco Duskywing
- Thanaos zarucco Lucas, 1857; ; TL: Cuba
- Nisoniades terentius Scudder & Burgess, 1870; Proc. Bost. Soc. Nat. Hist. 13: 292; TL: Florida
  - = ; [NL4A], #765
- Nisoniades ovidius Scudder & Burgess, 1870; ; TL: Florida
  - = ; [NL4A], #765
- Nisoniades naevius Lintner, 1881; Papilio 1 (5): 69; TL: Florida
  - = ; [NL4A], #765
- Antigonus diogenes Plötz, 1884; ; TL: Cuba
  - = ; [NL4A], #765
- Erynnis zarucco, Moth Photographers Group [MPG]
- Erynnis zarucco, Butterflies of America [BOA]
- Erynnis zarucco, Butterflies and Moths of North America [BAMONA]
- Nisoniades jaruco; Gundlach, 1881, (missp.)
- Antigonus ferentius; Plötz, 1884, (missp.)
- Thanaos terentius; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- Thanaos naevius; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- Nisoniades jaracco; Wolcott, 1936, (missp.)
- Erynnis zaruco; Silva et al., 1968, (missp.)
- jarucco; Beattie, 1976, (missp.)
- Erynnis zarucco; [NACL], #3956; [BOW]: pl. 21, f. 35 (text); [Opler]
- Erynnis (Erynnides) zarucco; [NL4A], #765
Larva on Robinia [NAB]

Subgenus ?

Erynnis lucilius (Scudder & Burgess, 1870) S.USA - Mexico. See [maps] Columbine Duskywing
- lucilius (Scudder & Burgess, 1870) (Nisoniades); Proc. Bost. Soc. Nat. Hist. 13: 287
- ?lucilius (Lintner, 1872); 23 Rept. N.Y. Sta. Cab. Nat. Hist.: 162
- Erynnis lucilius, Moth Photographers Group [MPG]
- Erynnis lucilius, Butterflies of America [BOA]
- Erynnis lucilius, Butterflies and Moths of North America [BAMONA]
- ?Thanaos lucilius Lintner, 1872; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- Erynnis lucilius; [NACL], #3958; [BOW]: pl. 21, f. 38 (text); [Opler]
Larva on Aquilegia canadensis [NAB]
Erynnis baptisiae (Forbes, 1936) E.USA. See [maps] Wild Indigo Duskywing
- Erynnis baptisiae, Moth Photographers Group [MPG]
- Erynnis baptisiae, Butterflies of America [BOA]
- Erynnis baptisiae, Butterflies and Moths of North America [BAMONA]
- Erynnis baptisiae; [NACL], #3959; [BOW]: pl. 21, f. 38 (text); [Opler]
Larva on Rubus chamaemorus [NAB]
Erynnis persius (Scudder, 1863) USA, Canada. See [maps] Persius Duskywing Hairy Dusky Wing
- persius (Scudder, 1863); Proc. Essex Inst. 3: 170
- Erynnis persius, Moth Photographers Group [MPG]
- Erynnis persius persius, Butterflies of America [BOA]
- Erynnis persius, Butterflies and Moths of North America [BAMONA]
- Thanaos persius; Dyar, 1903, Bull. U.S. nat. Mus. 52: 60
- Erynnis persius; [NACL], #3961; [BOW]: pl. 21, f. 38; [BOC], 55; [Opler]
Larva on Lupinus , Thermopsis [NAB], Lupinus latifolius, L. sericeus, Thermopsis , Astragalus , Lotus [BOC]
- E. p. persius
- E. p. borealis (Cary, 1906)
  - Thanaus propertius var. borealis Cary, 1906; Proc. U.S. Nat. Mus. 31 (1488): 455; TL: N.Nahanni R., Mckenzie Dist., NW Terr.
  - Erynnis persius borealis, Butterflies of America [BOA]
  - Erynnis persius borealis; [NACL], #3961a; [BOC], 55
- E. p. avinoffi (Holland, 1930)
  - Erynnis persius avinoffi, Butterflies of America [BOA]
  - Erynnis persius avinoffi; [NACL], #3961b
- E. p. fredericki Freeman, 1943
  - Erynnis persius fredericki Freeman, 1943; Ent. News 54 (3): (72-77) ; TL: near Lead, Spearfish Canyon, [Lawrence Co.,] South Dakota
  - Erynnis persius fredericki, Butterflies of America [BOA]
  - Erynnis persius fredericki; [NACL], #3961c
  - Erynnis persius frederecki[sic]; [BOC], 55

Unknown or unplaced taxa

Thanaos tages f. posticeprivata Stauder, 1924; Mitt. Münch. Ent. Ges. 14 (1-5): 66

12.3.2019 (24)

References:

[BAMONA] Opler, Pavulaan, Stanford & Pogue
Butterflies and Moths of North America; Mountain Prairie Information Node
[BCA] electronic
BIOLOGIA CENTRALI-AMERICANA
[BOA] Interactive Listing of American Butterflies
Listado Interactivo de la Mariposas Americanas; Butterflies of America
[Bruce Walsh]
Bruce Walsh; e-mail: Bruce Walsh; The Moths of Southeastern Arizona; Butterflies of Southeastern Arizona
[CiNii]
http://ci.nii.ac.jp/
[HBN] Francis Hemming, 1937
HÜBNER, A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements thereto by Carl Geyer, Gottfried Franz von Frölich and Gottlieb August Wilhelm Herrich-Schäffer, Vol 1 & 2
[Heiner Ziegler]
Switzerland / Schweiz; www.pieris.ch
[Kimmo Silvonen] Espoo, Finland
Kimmo's Lep Site
[LAMAS-NEO] Gerardo Lamas
Bibliography of butterflies; An Annocated Bibliography of the Neotropical Butterflies and Skipper (Lepididoptera: Papilionoidea and Hesperioidea); Revised Electronic Edition (2007)
[LEPIFORUM]
Lepiforum; Bestimmung von Schmetterlingen (Lepidoptera) und ihren Präimaginalstadien
[MBE] Moths and Butteflies of Europe
Paolo Mazzei, Diego Reggianti and Ilaria Pimpinelli; http://www.leps.it/
[MPG]
North American Moth Photographers Group
[NHM] Natural History Museum
The Global Lepidoptera Names Index; Butterflies and Moths of the World; Generic Names and their Type-species
[Oleg Kosterin] Institue of Cytology & Genetics
Siberian Division of Russian Academy of Sciences; Novosibirsk; e-mail: Oleg Kosterin; http://pisum.bionet.nsc.ru/kosterin/
[Opler] Opler & Warren, 2003 (April)
Butterflies of North America. 2. Scientific Names List for Butterfly Species of North America, north of Mexico.
[SVF]
Svenska fjärilar; Naturhistoriska riksmuseet; (Sweden)
[SZMN] Lepidoptera
Siberian Zoological Museum
[Simon Coombes] 1 Park Street, Plymouth, PL3 4BL, UK
01752 607854; e-mail: Simon Coombes; http://www.butterfly-guide.co.uk
[ZOBODAT] Oberösterreichische Landesmuseum
OÖ. Landesmuseum; ZOBODAT - Literatur
[maps]
Warning! The maps are automatically generated from the textual information, and the process does not always produce acceptable result; See about maps for more info.

Some related literature:

Bell, 1927
Description of a new species of Erynnis (Thanaos Auct.) (Lepidoptera-Rhopalocera-Hesperiidae) J. N.Y. Ent. Soc. 35 (3) : 261-263
Benyamini, Ugarte, Shapiro, Mielke, Pyrcz & Bálint, 2014
An updated list of the butterflies of Chile (Lepidoptera, Papilionoidea and Hesperioidea) including distribution, flight period and conservation status. Part I, Comprising the families: Papilionidae, Pieridae, Nympalidae (in part) and Hesperiidae. Describing a new species of Hypsochila (Pieridae) and a new subspecies of Yramea modesta (Nymphalidae) Bol. Mus. Nac. Hist. Nat. Chile 63 : 9-31
Boisduval, [1834]
Icones historique des Lépidoptères nouveaux ou peu connus. Collection, avec Figures coloritées, des Papillons d'Europe nouvellement découverts; ouvrage formant le complément de tous les auteurs iconographes Icon. hist. Lépid. Europe 1 (1-2): 1-24, pl. 1-4 (1832), (3-4): 25-40, pl. 5-8 (1832), (5-6): 41-56, pl. 9-12 (1832), (7-8): 57-72, pl. 13-16 (1832), (9-10): 73-104, pl. 17-20 (1833), (11-12): 105-120, pl. 21-24 (1833), (13-14): 121-144, pl. 25-28 (1833), (15-16): 145-160, pl. 29-32 (1833), (17-18): 161-184, pl. 33-36 (1833), (19-20): 185-208, pl. 37-40 (1834), (21-22): 209-224, pl. 41-44 (1834), (23-24): 225-248, pl. 45-47 (1834), (25-28): 249-252 (1834)
Boisduval, 1852
Lépidoptères de la Californie Ann. Soc. ent. Fr. (2) 10 (2) : 275-324
Boisduval & Le Conte, [1834]
Histoire générale et iconographie des lépidoptères et des chenilles de l'Amérique septentrionale Hist. Lép. Am. Sept. (1): 1-16. pl. 1-3 (1829), (2): 17-24, pl. 4-6 (1829), (3/4): 25-40, pl. 7-12 ([1830]), (5/6): 41-56, pl. 13-18 ([1830]), (7/8): 57-80, pl. 19-24 ([1830]), (9/10): 81-100, pl. 25-30 (1833), (11-22): 101-196, pl. 31-65 ([1835]), (23-26): 197-228, pl. 66-78 ([1837])
Bremer, 1861
Neue Lepidopteren aus Ost-Sibirien und dem Amur-Lande gesammelt von Radde und Maack, bescrieben von Otto Bremer Bull. Acad. Imp. Sci. St. Petersb. 3 : 461-496, Mélanges biol. St.-Pétersb. 3 (5-6): 538-589
Burns, 1960
A new species of oak-eating Erynnis (Lepidoptera: Hesperiidae) Wasmann J. Biol. 18 (1): 147-160, f. 1-7
Butler, 1866
A list of the diurnal Lepidoptera recently collected by Mr. Whitely in Hakodadi (North Japan) J. Linn. Soc. Zool. Lond. 9 (34) : 50-58
Bytinski-Salz & Brandt, 1937
New Lepidoptera from Iran Ent. Rec. J. Var. (Suppl.) 49 : 1-9, : 9-15
Cary, 1906
On the Diurnal Lepidoptera of the Athabaska and Mackenzie Region, British America Proc. U.S. Nat. Mus. 31 (1488) : 425-457
[MRS]; Chou, 1994
Monografia Rhopalocerorum Sinensium 1-2: 1-854
Dunning & Pickard, 1858
An Accentuated List of the British Lepidoptera Accent. List Brit. Lep.: : 1-118
Dyar, 1903
A List of North American Lepidoptera and Key to the Literature of this Order of Insects Bull. U.S. nat. Mus. 52 : 1-723
Edwards, 1882
Two new species of Thanaos Papilio 2 (9-10) : 179-180
Evans, 1949
A catalogue of the Hesperiidae from Europe, Asia and Australia in the British Museum Cat. Hesp. Europe Asia Australia Brit. Mus. : 1-502, pl. 1-53
Fabricius, 1807
Die neueste Gattungs-Eintheilung der Schmetterlinge aus den Linnéischen Gattungen Papilio und Sphinx Mag. f. Insektenk. 6 : 277-295
Freeman, 1943
New Hesperioidea, with Notes on Some Others from the United States (Lepidotera, Rhopalocera) Ent. News 54 (3) : 72-77
Godman & Salvin, [1899]; Godman in Godman & Salvin, [1899]
Biologia Centrali-Americana. Rhopalocera. (1887-1901) Biol. centr.-amer., Lep. Rhop. 2: 1-782, 3: pl. 1-112
Grum-Grshimailo, 1891
Lepidoptera nova in Asia centrali novissime lecta et descripta Horae Soc. ent. Ross. 25 (3-4) : 445-465
[H&R]; Higgins & Riley, 1970
A Field Guide to the Butterflies of Britain and Europe
[NACL]; Hodges, 1983
Check List of the Lepidoptera of America North of Mexico Check List Lep. Am. N of Mexico
Hübner, 1822
Systematisch-alphabetisches Verzeichniss aller bisher bey den Fürbildungen zur Sammlung europäischer Schmetterlinge angegebenen Gattungsbenennungen; mit Vormerkung auch augsburgischer Gattungen, von Jacob Hübner Syst.-alph. Verz. : I-III, IV-VI, 1-81
Korb & Bolshakov, 2011
[A catalogue of butterflies (Lepidoptera: Papilioformes) of the former USSR. Second edition, reformatted and updated] (in Russian) Eversmannia Suppl. 2 : 1-123
Kudrna, 1974
An annotated List of Japanese Butterflies Atalanta 5 : 92-120
[NL4A]; Lamas, 2004
Checklist: Part 4A, Hesperioidea - Papilionoidea Atlas of Neotropical Lepidoptera
Leech, 1891
New Species of Rhopalocera from Western China Entomologist 24 (Suppl.) : 57-61, 66-68
[RFEL]; Leley, 2016
Annotated catalogue of the insects of Russian Far East. Volume II. Lepidoptera Cat. ins. Russian Far East. 2 : 1-812
[BOW]; Lewis, 1974
Butterflies of the World
Linnaeus, 1758
Systema Naturae per Regna Tria Naturae, Secundum Clases, Ordines, Genera, Species, cum Characteribus, Differentiis, Symonymis, Locis. Tomis I. 10th Edition Syst. Nat. (Edn 10) 1 : 1-338, : 339-824
Lintner, 1881
On some species of Nisoniades Papilio 1 (5) : 69-74
Mabille, 1883
Description d'hespériens Bull. Soc. Ent. Belg. 27 : li-lxxviii
Murayama & Yoshisaka, 1959
Die neu aus Formosa entdeckte Art Erynnis montanus BREMER [in Japanese] Tyô to Ga 10 (1): 14
Nordmann, 1851
Neue Schmetterlinge Russlands Bull. Soc. imp. Nat. Moscou 24 : 439-446, pl. 11-12
[NAB]; Pyle, 1981
Field Guide to North American Butterflies, National Audubon Society
[BOC]; Pyle, 2002
The Butterflies of Cascadia
Schrank, 1801
Fauna Boica. Durchgedachte Geschichte der in Baiern einheimischen und zahmen Thiere Fauna Boica 2 (1) : 1-374
Scudder, 1863
A List of the Butterflies of New England Proc. Essex Inst. 3 : 161-179
Stauder, 1924
Neue Palaearktenformen. II Mitt. Münch. Ent. Ges. 14 (1-5) : 59-66
Tutt, 1906
A natural history of the British Lepidoptera. A text-book for students and collectors Nat. Hist. Br. Lepid. 8 : 1-479, pl. 1-20
[BRU]; Tuzov, Bogdanov, Devyatkin, Kaabak, Korolev, Murzin, Samodurov, Tarasov, 1997
Guide to the Butterflies of Russia and adjacent territories: Hesperiidae, Papilionidae, Pieridae, Satyridae Butts. Russia adj. terr. 1
Yakovlev, 2012
Checklist of Butterflies (Papilionoidea) of the Mongolian Altai Mountains, including descriptions of new taxa Nota lepid. 35 (1) : 51-96

If you have corrections, comments or information to add into these pages, just send mail to Markku Savela
Keep in mind that the taxonomic information is copied from various sources, and may include many inaccuracies. Expert help is welcome.

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